Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Cardioglossa gratiosa Amiet, 1972

Class: Amphibia > Order: Anura > Family: Arthroleptidae > Subfamily: Arthroleptinae > Genus: Cardioglossa > Species: Cardioglossa gratiosa

Cardioglossa gratiosa Amiet, 1972, Biol. Gabonica, 8: 221. Holotype: MHNG 1253.85 (formerly JLA 71.070) according to Amiet, 1985, in Frost (ed.), Amph. Species World: 16; and Schätti, Perret, and Mariaux, 2002, Cat. Comm. Types Amph. Rept. Mus. Hist. Nat. Geneve, Vers. 4.0 (Jan. 2002): 7. Type locality: "Ongot, env. 750 m", Cameroon.

Cardioglossa gratiosa peternageli Böhme and Schneider, 1987, Bonn. Zool. Beitr., 38: 249. Holotype: ZFMK 40805, by original designation. Type locality: "Minim," Cameroon.

Cardioglossa gratiosa gratiosa — Böhme and Schneider, 1987, Bonn. Zool. Beitr., 38: 249.

Cardioglossa peternageli — Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 38. Unsubstantiated rank elevation.

Astylosternus gratiosus — Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 222, by implication. See comment under Cardioglossa.

Common Names

Ongot Long-fingered Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 37).

Nagel's Long-fingered Frog (Cardioglossa gratiosa peternageli: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 38).

Pleasing Long-fingered Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 242).

Distribution

Widely distributed in Cameroon south of 7° 5´ N, except in the littoral plain and mountains south through Equatorial Guinea to southern Gabon; north-central Dem. Rep. Congo; presumably in southwestern Central African Republic and northern Rep. Congo.

Geographic Occurrence

Natural Resident: Cameroon, Congo, Democratic Republic of the, Equatorial Guinea, Gabon

Likely/Controversially Present: Central African Republic, Congo, Republic of the

Comment

Closely related to Cardioglossa nigromaculata according to Amiet, 1985, in Frost (ed.), Amph. Species World: 17. This was rejected by Blackburn, 2008, Zool. J. Linn. Soc., 154: 624. Penner and Rödel, 2007, in Butynski and McCullough (eds.), RAP Bull. Biol. Assessment, 46: 37-41, provided a record for north-central Dem. Rep. Congo. Dewynter and Frétey, 2019, Cah. Fondation Biotope, 27: 9, summarized the literature for Gabon and provided a photograph. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 242–243, provided a brief account, photograph, and range map. See brief account, range map, and photographs for Equatorial Guinea in Sánchez-Vialas, Calvo-Revuelta, Castroviejo-Fisher, and De la Riva, 2020, Proc. California Acad. Sci., Ser. 4, 66: 137–230. Kako-Wanzalire, Mongo, Ilonga, Mapoli, Mbumba, Neema, Tungaluna, Itoka, and Bogaert, 2021, Tropicultura, 39 (1: 1709): 1–19, briefly discussed habitat preference in north-central Dem. Rep. Congo.

External links:

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.