Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptopelis bufonides Schiøtz, 1967

Class: Amphibia > Order: Anura > Family: Arthroleptidae > Subfamily: Leptopelinae > Genus: Leptopelis > Species: Leptopelis bufonides

Leptopelis bufonides Schiøtz, 1967, Spolia Zool. Mus. Haun., 25: 27. Holotype: ZMUC R073369, by original designation. Type locality: "a few miles west of Bolgatanga, Ghana, near the road to Navrongo".

Common Names

Savannah Forest Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 69).

Ground Tree Frog (Emms, Jambang, Bah, Mankali, Rödel, and Barnett, 2005, Herpetol. Bull., London, 94: 14).

Toad-like Tree Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 212).

Distribution

Savannas of Senegal and central-northern Guinea to southern Mauritania, northern Cameroon, Burkina Faso, and southern Chad; expected in Ivory Coast, southern Mali, and southern Niger.

Geographic Occurrence

Natural Resident: Benin, Burkina Faso, Cameroon, Chad, Gambia, Ghana, Guinea, Guinea-Bissau, Mauritania, Nigeria, Senegal, Togo

Likely/Controversially Present: Ivory Coast, Mali, Niger

Comment

See Amiet, 1973 "1972", Ann. Fac. Sci. Cameroun, 12: 71, and Amiet and Schiøtz, 1974, Ann. Fac. Sci. Cameroun, 17: 139, for distribution, ecology, and sonogram in Cameroon. Schiøtz, 1999, Treefrogs Afr.: 286, suggested that this form may be conspecific with Leptopelis bocagei. Rödel, 2000, Herpetofauna W. Afr., 1: 192–193, provided an account. Gambia record by Barnett, Emms, and Santoni, 2001, Herpetol. Bull., London, 77: 8. Padial and De la Riva, 2004, Rev. Esp. Herpetol., 18: 96, suggested that this species will be found in southern Mauritania. Wanger, 2005, Salamandra, 41: 27–33, provided Gambia records. Nago, Grell, Sinsin, and Rödel, 2006, Salamandra, 42: 93–108, provided a record for Benin. Emms, Jambang, Bah, Mankali, Rödel, and Barnett, 2005, Herpetol. Bull., London, 94: 6–16, provided records for Gambia. Hillers, Loua, and Rödel, 2008, Salamandra, 44: 113–122, provided the first record for Guinea. Auliya, Wagner, and Böhme, 2012, Bonn Zool. Bull., 61: 255–281, reported the species from the Bijagos Archipelago, Guinea-Bissau. See account for Cameroon by Amiet, 2012, Rainettes Cameroun: 487–491. Schiøtz, 1999, Treefrogs Afr.: 285–288, provided a brief account and map. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 212–213, provided a brief account, photograph, and range map. Ayoro, Segniagbeto, Hema, Penner, Oueda, Dubois, Rödel, Kabré, and Ohler, 2020, Zoosystema, 42: 547–582, discussed records, identification, and habitat in Burkina Faso. Sampaio, Velo-Antón, Martínez-Freiría, Sánchez-Vialas, Pleguezuelos, Géniez, Crochet, and Brito, 2021, Conserv. Genetics, 22: 233–248, reported the species from Mauritania.

External links:

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For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.