Leptopelis calcaratus (Boulenger, 1906)

Class: Amphibia > Order: Anura > Family: Arthroleptidae > Subfamily: Leptopelinae > Genus: Leptopelis > Species: Leptopelis calcaratus

Hylambates calcaratus Boulenger, 1906, Ann. Mag. Nat. Hist., Ser. 7, 17: 322. Syntypes: BMNH, by original designation. Type locality: "Efulen", Cameroon, and "Cape St. John and the Rio Benito District (Spanish West Africa [= Equatorial Guinea]".

Leptopelis calcaratusNoble, 1924, Bull. Am. Mus. Nat. Hist., 49: 234, 237, 332.

Leptopelis (Leptopelis) calcaratusLaurent, 1941, Rev. Zool. Bot. Afr., 35: 94.

Leptopelis calcaratus calcaratusLaurent, 1973, Ann. Mus. R. Afr. Cent., Tervuren, Ser. Octavo, Sci. Zool., 202: 50.

Leptopelis calcaratus meridionalis Laurent, 1973, Ann. Mus. R. Afr. Cent., Tervuren, Ser. Octavo, Sci. Zool., 202: 50. Holotype: MRAC 115147, by original designation. Type locality: "Omaniundu, Terr. de Lodja, Sankuru, Zaïre".

English Names

Efulen Forest Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 69).

Spurred Tree Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 216).


Dense forest from southeastern Nigeria through southern Cameroon to eastern Dem. Rep. Congo, and western Uganda, south through much of Gabon and northern Rep. Congo; Bioko; reported from Uigé Province of northwestern Angola.


De la Riva, 1994, Rev. Esp. Herpetol., 8: 130, provided a record for Equatorial Guinea. Lasso, Rial, Castroviejo, and De la Riva, 2002, Graellsia, 58: 21-34, provided notes on ecological distribution in Equatorial Guinea. Schiøtz, 1963, Vidensk. Medd. Dansk Naturhist. Foren., 125: 87, provided records for Nigeria. Frétey, Barriere, and Ohler, 2006, Herpetol. Rev., 37: 358, provided a record for the Central African Republic and delimited the range. Jackson and Blackburn, 2007, Salamandra, 43: 149-164, provided a record for Nouabale-Ndoki National Park, Rep. Congo. See account for Cameroon by Amiet, 2012, Rainettes Cameroun: 492-498. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 110, reported on comparative tadpole morphology.  Schiøtz, 1999, Treefrogs Afr.: 260–262, provided an account and map. Barej, Pfalzgraff, Hirschfeld, Liedtke, Penner, Gonwouo, Dahmen, Grözinger, Schmitz, and Rödel, 2015, Amph. Rept. Conserv., 9(2, Spec. Section): 56–84, reported on larval morphology. Jongsma, Tobi, Dixon-MacCallum, Bamba-Kaya, Yoga, Mbega, Mve Beh, Emrich, and Blackburn, 2017, Amph. Rept. Conserv., 11 (1: e144): 6, provided a record for southeastern Gabon and briefly discussed habitat and range. Dewynter and Frétey, 2019, Cah. Fondation Biotope, 27: 13, summarized the literature for Gabon. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 216–217, provided a brief account, photograph, and range map. Mali, Banda, Kusamba, Babangenge, Sebe, Lokasola, Ewango, Tungaluna, and Akaibe, 2019, Am. J. Zool., 2: 38–43, reported the species from the Okapi Wildlife Reserve, northeastern Dem. Rep. Congo. See brief account, range map, and photographs for Equatorial Guinea in Sánchez-Vialas, Calvo-Revuelta, Castroviejo-Fisher, and De la Riva, 2020, Proc. California Acad. Sci., Ser. 4, 66: 137–230, who suggested that the Bioko I. population might be a different species. Ernst, Lautenschläger, Branquima, and Hölting, 2020, Zoosyst. Evol., 96: 249, noted the species to occur in Uigé Province of northwestern Angola. Greenbaum, Hughes, and Behangana, 2021, Herpetol. Rev., 52: 342, provided a record from Buhoma, Bwindi Impenetrable National Park, Western Region, Uganda, 1523 m. 

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