Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptopelis gramineus (Boulenger, 1898)

Class: Amphibia > Order: Anura > Family: Arthroleptidae > Subfamily: Leptopelinae > Genus: Leptopelis > Species: Leptopelis gramineus

Megalixalus gramineus Boulenger, 1898, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 2, 18: 721. Syntypes: BMNH and MSNG (several specimens), including BMNH 1898.1.28.20–21 (reregistered 1947.2.10.19–20) according to museum records; MSNG 28564A designated lectotype by Capocaccia, 1957, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 3, 69: 217. Type locality: "between Badditù and Dimè", Ethiopia. Type locality restricted (via an extended discussion) to "Ethiopia, Gamu Gofa, 6°29′N, 36°59′E" by Tiutenko and Zinenko, 2021, ZooKeys, 1023: 119.

Leptopelis gramineus — Parker, 1930, Proc. Zool. Soc. London, 1930: 1; Tiutenko and Zinenko, 2021, ZooKeys, 1023: 127.

Megalixalus gramineus — Ahl, 1931, Das Tierreich, 55: 432.

Common Names

Badditu Forest Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 69). Inappropriate name due to pointing to a location outside of known range ( Tiutenko and Zinenko, 2021, ZooKeys, 1023: 124).

Ethiopian Burrowing Tree Frog (Largen and Spawls, 2010, Amph. Rept. Ethiopia Eritrea: 126).

Distribution

Ethiopian Highlands, 1900–3900 m elevation, west of the Rift Valley.

Geographic Occurrence

Natural Resident: Ethiopia

Endemic: Ethiopia

Comment

Largen, 2001, Tropical Zool., 14: 350-351, discussed distribution. See account, photograph, and map for Ethiopia by Largen and Spawls, 2010, Amph. Rept. Ethiopia Eritrea: 135-136. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 110–111, reported on comparative tadpole morphology. Schiøtz, 1999, Treefrogs Afr.: 288–289, provided a brief account and map. Freilich, Anadón, Bukala, Calderon, Chakraborty, and Boissinot, 2016, BMC Evol. Biol., 16 (206): 1–19, reported on the molecular phylogeography from both sides of the Great Rift in montane Ethiopia. Reyes-Velasco, Manthey, Freilich, and Boissinot, 2018, Mol. Ecol., 27: 2256–2270, discussed the molecular phylogenetics of the species of the Ethiopian highlands, showing nominal Leptopelis gramineus to be a species complex, with Leptopelis susanae being imbedded within the group. Stand by for the usual suspects to swoop in and name these out from under these authors (DRF). Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 224–225, provided a brief account, photograph, and range map. Tiutenko and Zinenko, 2021, ZooKeys, 1023: 119–150, showed that nominal Leptopelis gramineus east and west of the Great Rift are different species, with the eastern population named Leptopelis diffidens. These authors also noted that nominal Leptopelis gramineus west of the Great Rift is composed of two lineages of which the northerly one is unnamed.

External links:

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For access to general information see Wikipedia
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For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.