Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptopelis kivuensis Ahl, 1929

Class: Amphibia > Order: Anura > Family: Arthroleptidae > Subfamily: Leptopelinae > Genus: Leptopelis > Species: Leptopelis kivuensis

Leptopelis kivuensis Ahl, 1929, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1929: 206. Holotype: ZMB unnumbered, according to the original publication. Type locality: "Kisenji, südlich des Kivu-Sees", Rwanda.

Leptopelis graueri Ahl, 1929, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1929: 213. Holotype: ZMB unnumbered, according to the original publication. Type locality: "70 km westlich des Südendes des Albert-Edward-See's". Synonymy by Laurent, 1972, Explor. Parc Natl. Virunga, Ser. 2, 22: 47. Formerly considered a synonym of Leptopelis karissimbensis by De Witte, 1941, Explor. Parc Natl. Albert, Miss. G.F. de Witte (1933–1935), 33: 94.

Leptopelis rugegensis Ahl, 1929, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1929: 218. Syntypes: ZMB (4 specimens) unnumbered, according to the original publication; MCZ 17531 (on exchange from ZMB) is a syntype according to Barbour and Loveridge, 1946, Bull. Mus. Comp. Zool., 96: 141, status unknown. Type locality: "Rugege-Wald, Deutsch-Ost-Afrika". Synonymy by Laurent, 1972, Explor. Parc Natl. Virunga, Ser. 2, 22: 47. Formerly considered a synonym of Leptopelis karissimbensis by Loveridge, 1936, Field Mus. Nat. Hist. Publ., Zool. Ser., 22: 102.

Common Names

Kisenyi Forest Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 69).

Kivu Tree Frog (Channing and Howell, 2006, Amph. E. Afr.: 204).

Distribution

On the east side of the Albertine Rift in the highlands of western Rwanda, Burundi; west of the Albertine Rift from adjacent North and South Kivu province, Dem. Rep. Congo; unnamed species (see comment) noted in a) southwestern Rwanda and northwestern Burundi; b) northwestern Rwanda and southwestern Uganda; c) and south-central Burundi; and d) eastern North Kivu Province, Dem. Rep. Congo.

Geographic Occurrence

Natural Resident: Burundi, Congo, Democratic Republic of the, Rwanda, Uganda

Comment

Redescribed as new by Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 26. See account by Laurent, 1972, Explor. Parc Natl. Virunga, Ser. 2, 22: 47, who removed Leptopelis kivuensis from the synonymy of Leptopelis karissimbensis, where it had been placed by De Witte, 1941, Explor. Parc Natl. Albert, Miss. G.F. de Witte (1933–1935), 33: 94, and suggested a widespread distribution. Schiøtz, 1999, Treefrogs Afr.: 273, provided a brief account and map and suggested the possibility of considerable confusion with Leptopelis karissimbensis, and gave a much restricted distribution for Leptopelis kivuensis (montane forest from eastern Dem. Rep. Congo and southwestern Uganda). The status of populations in the larger stated distribution is unclear. Channing and Howell, 2006, Amph. E. Afr.: 204-205, provided an account. See statement of geographic range, habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 618. Roelke, Greenbaum, Chifundera, Aristote, and Smith, 2011, J. Herpetol., 45: 343-351, discussed the systematics and range of this species and the very similar Leptopelis karissimbensis. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 112, reported on comparative tadpole morphology. Portillo, Greenbaum, Menegon, Chifundera, and Dehling, 2015, Mol. Phylogenet. Evol., 82: 75–86, discussed the phylogeography of the species, noting that nominal Leptopelis kivuensis is a complex, with at least two unnamed associates on the east side of the Albertine Rift in Uganda, Rwanda, Dem. Rep. Congo, and Uganda. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 222–223, provided a brief account, photograph, and range map. Dehling and Sinsch, 2023, Diversity, 15 (512): 1–81, discussed the range, identification, natural history, advertisement call, and conservation status. Behangana, Byaruhanga, Magala, Katumba, Kagurusi, Dendi, and Luiselli, 2023, Wetlands, 43(88): 1–13, reported on presence in the wetlands of southwestern Uganda.

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For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.