Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptopelis modestus (Werner, 1898)

Class: Amphibia > Order: Anura > Family: Arthroleptidae > Subfamily: Leptopelinae > Genus: Leptopelis > Species: Leptopelis modestus

Hylambates rufus var. modesta Werner, 1898, Verh. Zool. Bot. Ges. Wien, 48: 197. Syntypes: ZMB 114112 (2 specimens) (Cameroon), 20046 (2 specimens) (South Cameroon), 28708 (Buéa, Cameroon), and MCZ 17621 (on exchange from ZMB, according to Barbour and Loveridge, 1946, Bull. Mus. Comp. Zool., 96: 125); ZMB 28708 designated lectotype by Perret, 1962, Rev. Zool. Bot. Afr., 65: 237. Type locality: "Kamerun"; restricted to "Buéa, Cameroun" by lectotype designation.

Hylambates rufus var. ventrimaculata Werner, 1898, Verh. Zool. Bot. Ges. Wien, 48: 198. Types: ZMB, by original designation. Type locality: "Kamerun". Synonymy by Ahl, 1929, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1929: 215.

Leptopelis modestus — Ahl, 1929, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1929: 214.

Leptopelis modestus modestus — Laurent, 1973, Ann. Mus. R. Afr. Cent., Tervuren, Ser. Octavo, Sci. Zool., 202: 36.

Common Names

Modest Forest Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 69).

Plain Tree Frog (Channing and Howell, 2006, Amph. E. Afr.: 205).

Distribution

Medium and high-altitude forests in southeastern Nigeria and Cameroon; eastern Dem. Rep. Congo; Bioko I. (Equatorial Guinea).

Geographic Occurrence

Natural Resident: Cameroon, Congo, Democratic Republic of the, Equatorial Guinea, Nigeria

Comment

See brief account in Laurent, 1972, Explor. Parc Natl. Virunga, Ser. 2, 22: 46-47. Schiøtz, 1999, Treefrogs Afr.: 269, discussed unnamed species masquerading under this name: one found at high altitudes in Cameroon and on the Obudu Plateau in Nigeria, the other likely represented by the East African population. Channing and Howell, 2006, Amph. E. Afr.: 205-206, provided an account. See account for Cameroon by Amiet, 2012, Rainettes Cameroun: 513-521. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 113–114, reported on comparative tadpole morphology. Schiøtz, 1999, Treefrogs Afr.: 269–271, provided an account and map in the sense of including unnamed and now-named (e.g., Leptopelis mtoewaate, Leptopelis mackayi) species. Barej, Pfalzgraff, Hirschfeld, Liedtke, Penner, Gonwouo, Dahmen, Grözinger, Schmitz, and Rödel, 2015, Amph. Rept. Conserv., 9(2, Spec. Section): 56–84, reported on larval morphology. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 220–221, provided a brief account, photograph, and range map. See brief account, range map, and photographs for Equatorial Guinea in Sánchez-Vialas, Calvo-Revuelta, Castroviejo-Fisher, and De la Riva, 2020, Proc. California Acad. Sci., Ser. 4, 66: 137–230.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.