Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptopelis vermiculatus (Boulenger, 1909)

Class: Amphibia > Order: Anura > Family: Arthroleptidae > Subfamily: Leptopelinae > Genus: Leptopelis > Species: Leptopelis vermiculatus

Hylambates vermiculatus Boulenger, 1909, Ann. Mag. Nat. Hist., Ser. 8, 4: 497. Holotype: BMNH, by original designation; identified as BMNH 1947.2.20.11 (formerly 1909.10.19.20), by Gvoždík, Tillack, Menegon, and Loader, 2014, Zootaxa, 3793: 167. Type locality: "Amani", Usambara Mountains, Tanzania.

Leptopelis vermiculatus — Barbour and Loveridge, 1928, Mem. Mus. Comp. Zool., 50: 238.

Leptopelis signifer Ahl, 1929, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1929: 216. Syntypes: ZMB (3 specimens) unnumbered, according to the original publication; one of these, MCZ 17530 (on exchange from ZMB) is likely a paratype, not a syntype (see Barbour and Loveridge, 1946, Bull. Mus. Comp. Zool., 96: 141); ZMB 11773A designated lectotype by Gvoždík, Tillack, Menegon, and Loader, 2014, Zootaxa, 3793: 181. Type locality: "Derema", Tanzania, this rendered as "'Derema', East Usambara Mountains, Tanzania, approx. 5.05°S 38.66°E" by Gvoždík, Tillack, Menegon, and Loader, 2014, Zootaxa, 3793: 181. De Witte, 1941, Explor. Parc Natl. Albert, Miss. G.F. de Witte (1933–1935), 33: 102, reported the syntypes ("cotypes") as being from "Urwald zwischen Beni und Mboga". Synonymy by Loveridge, 1933, Bull. Mus. Comp. Zool., 74: 346, and Schiøtz, 1975, Treefrogs E. Afr.: 39. Formerly considered a junior synonym of Leptopelis christyi by Laurent, 1972, Explor. Parc Natl. Virunga, Ser. 2, 22: 50.

Common Names

Amani Forest Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 69).

Vermiculated Tree Frog (Channing and Howell, 2006, Amph. E. Afr.: 210).

Distribution

Forests of the East and West Usambara, Nguu, Nguru, Udzungwa, Mahenge, and Poroto mountains, as well as Mount Rungwe, southern Tanzania, 900–1800 m elevation.

Geographic Occurrence

Natural Resident: Tanzania

Endemic: Tanzania

Comment

Schiøtz, 1975, Treefrogs E. Afr.: 42, noted that this species is easily confused with Leptopelis flavomaculatus. Schiøtz, 1999, Treefrogs Afr.: 280–281, and Channing and Howell, 2006, Amph. E. Afr.: 210–211, provided accounts. Loader, Poynton, and Mariaux, 2004, Afr. Zool., 39: 71–76, provided a record for Mahenge Mountain in Tanzania and detailed the range. Pickersgill, 2007, Frog Search: 491–492, provided a brief account. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 291. Menegon, Doggart, and Owen, 2008, Acta Herpetol., Firenze, 3: 107–127, provided a record for the Nguru Mountains of eastern Tanzania. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 120–121, reported on comparative tadpole morphology. See taxonomic and descriptive comments by Gvoždík, Tillack, Menegon, and Loader, 2014, Zootaxa, 3793: 165–187. Harper, Measey, Patrick, Menegon, and Vonesh, 2010, Field Guide Amph. E. Arc Mts. Tanzania and Kenya: 76–77, provided a brief account for the Eastern Arc Mountains of Tanzania. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 226–227, provided a brief account, photograph, and range map.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.