Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptopelis viridis (Günther, 1869)

Class: Amphibia > Order: Anura > Family: Arthroleptidae > Subfamily: Leptopelinae > Genus: Leptopelis > Species: Leptopelis viridis

Hylambates viridis Günther, 1869 "1868", Proc. Zool. Soc. London, 1868: 487. Holotype: BMNH, by original designation; BMNH 1947.2.10.23 by museum records. Type locality: Not stated; given as "W. Africa" by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 134.

Hylambates hyloides Boulenger, 1906 "1905", Ann. Mus. Civ. Stor. Nat. Genova, Ser. 3, 2: 167. Syntypes: "Numerous specimens" in MSNG and BMNH, the BMNH specimens being 1906.3.30.151–155 (reregistered as 1947.2.20.12–16) according to museum records; MSNG 29944A designated lectotype by Capocaccia, 1957, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 3, 69: 218. Type locality: "Bolama, Portuguese Guinea [= Guinea-Bissau]". Synonymy by Parker, 1936, Zool. Meded., Leiden, 19: 97, and Perret, 1966, Zool. Jahrb., Jena, Abt. Syst., 93: 432. A. Schiøtz In Laurent, 1985, in Frost (ed.), Amph. Species World: 230, noted that the types of Leptopelis hyloides were of the biological species Leptopelis viridis, but the species that is commonly associated with the name Leptopelis hyloides was another, now named Leptopelis spiritusnoctis.

Leptopelis nanus Ahl, 1924, Arch. Naturgesch., Abt. A,, 90: 252. Holotype: ZMB unnumbered according to the original publication; now ZMB 26990 (M.-O. Rödel, personal commun.). Type locality: "Mangu [now Sansanne-Mango], Togo". Synonymy by Parker, 1936, Zool. Meded., Leiden, 19: 97; Perret, 1966, Zool. Jahrb., Jena, Abt. Syst., 93: 432. (Formerly placed in the synonymy of Leptopelus bocagii by Loveridge, 1929, Bull. U.S. Natl. Mus., 151: 121.)

Leptopelis togoensis Ahl, 1929, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1929: 191. Syntypes: ZMB (2 specimens) unnumbered according to the original publication; these now possibly lost (M.-O. Rödel, personal commun.). Type locality: "Bumpana (Panpamba), Togo". Subsequently Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 113, stated the type locality to only be "Togo". Synonymy by Parker, 1936, Zool. Meded., Leiden, 19: 97; Perret, 1966, Zool. Jahrb., Jena, Abt. Syst., 93: 432.

Leptopelis liberiensis Ahl, 1929, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1929: 196. Syntypes: ZMB (2 specimens) unnumbered according to the original publication, these now possibly lost (M.-O. Rödel, personal commun.). Type locality: "Liberia". Synonymy by Loveridge, 1941, Proc. U.S. Natl. Mus., 91: 128.

Leptopelis hyloides — Ahl, 1931, Das Tierreich, 55: 215.

Leptopelis viridis — Ahl, 1931, Das Tierreich, 55: 212; Parker, 1936, Zool. Meded., Leiden, 19: 95.

Leptopelis nanus — Ahl, 1931, Das Tierreich, 55: 216.

Leptopelis (Taphriomantis) viridis — Laurent, 1941, Rev. Zool. Bot. Afr., 35: 102.

Common Names

Green Tree Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 212; Spawls, Mazuch, and Mohammad, 2023, Handb. Amph. Rept. NE Afr.: 80).

Rusty Forest Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 69).

Savannah Tree Frog (Emms, Jambang, Bah, Mankali, Rödel, and Barnett, 2005, Herpetol. Bull., London, 94: 14).

Gbanga Forest Treefrog (Leptopelis hyloides [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 69).

Distribution

Savanna of West Africa from Senegal and southern Niger to northeastern Dem. Rep. Congo, southern South Sudan, Uganda, and likely into Chad, Mali, and Central African Republic. See comment.

Geographic Occurrence

Natural Resident: Benin, Burkina Faso, Cameroon, Congo, Democratic Republic of the, Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Liberia, Niger, Nigeria, Senegal, Sierra Leone, South Sudan, Togo, Uganda

Likely/Controversially Present: Central African Republic, Chad, Mali

Comment

Literature of this species is confusing because the type of Leptopelis hyloides is conspecific with Leptopelis viridis, although often applied to a forest species distinct from Leptopelis viridis. So, nomenclaturally, Leptopelis hyloides became a junior synonym of Leptopelis viridis, while the living populations previously identifed as Leptopelis hyloides were given a new name—Leptopelis spiritusnoctis. All older records of Leptopelis hyloides (other than the type) now refer to Leptopelis spiritusnoctis. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 121–122, reported on comparative tadpole morphology. Schiøtz, 1999, Treefrogs Afr.: 300–301, provided a brief account and map. Auliya, Wagner, and Böhme, 2012, Bonn Zool. Bull., 61: 255–281, reported the species from the Bijagos Archipelago, Guinea-Bissau. Barej, Pfalzgraff, Hirschfeld, Liedtke, Penner, Gonwouo, Dahmen, Grözinger, Schmitz, and Rödel, 2015, Amph. Rept. Conserv., 9(2, Spec. Section): 56–84, reported on larval morphology. Marques, Ceríaco, Blackburn, and Bauer, 2018, Proc. California Acad. Sci., Ser. 4, 65 (Suppl. II): 126–127, provided a map and brief account for Angola, suggesting its presence there. Rödel and Glos, 2019, Zoosyst. Evol., 95: 20, reported this species from the Foya Proposed Protected Area in western Liberia. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 212–213, provided a brief account, photograph, and range map. Ayoro, Segniagbeto, Hema, Penner, Oueda, Dubois, Rödel, Kabré, and Ohler, 2020, Zoosystema, 42: 547–582, discussed records, identification, and habitat in Burkina Faso. Kanga, Kouamé, Zogbass, Gongomin, Agoh, Kouamé, Konan, Adepo-Gourène, Gourène, and Rödel, 2021, Amph. Rept. Conserv., 15: 71–107, commented on conservation status, identification, range, and habitat on the Ivory Coast side of Mont Nimba. Gansa, Agadjihouèdé, and Hounkanrin, 2023, Afr. Zool., 58: 39–56, reported the species from the lower Ouémé Valley, southern Benin, and briefly descried habitat preference. Segniagbeto, Ohler, Rödel, Luiselli, and Dubois, 2024, Zoosystema, 46: 638–639, provided an account for Togo, discussing habitat, conservation status, and identification. Spawls, Mazuch, and Mohammad, 2023, Handb. Amph. Rept. NE Afr.: 80, provided an account addressing identification, natural history, conservation status, and range in South Sudan, including a dot map.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.