Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Craugastor campbelli (Smith, 2005)

Class: Amphibia > Order: Anura > Superfamily: Brachycephaloidea > Family: Craugastoridae > Genus: Craugastor > Species: Craugastor campbelli

Eleutherodactylus campbelli Smith, 2005, Herpetologica, 61: 287. Holotype: UTA A-33452, by original designation. Type locality: "Torres de Guatel, Cerro San Gil, Montañas del Mico, Municipio de Puerto Barrios, Departamento de Izabal, Guatemala, 962 m".

Eleutherodactylus nefrens Smith, 2005, Herpetologica, 61: 287. Holotype: UTA A-45279, by original designation. Type locality: "La Firmeza, Sierra de Caral, Municipio de Morales, Departamento de Izabal, Guatemala, 840 m". Synonymy by Streicher, Wiens, Jocqué, García-Vázquez, and Smith, 2023, J. Vert. Biol., 72(23072): 19.

Craugastor campbelli — Crawford and Smith, 2005, Mol. Phylogenet. Evol., 35: 551, by implication; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 360.

Craugastor cyanochthebius McCranie and Smith, 2006, Herpetologica, 62: 185. Holotype: UTA A-56748, by original designation. Type locality: "San Isidro, on the western slopes of Montaña del Cerro Azul, Sierra del Espíritu Santo, Departamento de Copán, Honduras, 1200 m elevation. . . . The locality is approximately 15° 07′ 38.2″ N, 88° 56′ 13.8″ W". Synonymy by Streicher, Wiens, Jocqué, García-Vázquez, and Smith, 2023, J. Vert. Biol., 72(23072): 19.

Craugastor (Hylactophryne) campbelli — Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 45.

Craugastor (Hylactophryne) cyanochthebius — Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 45.

Common Names

None noted.

Distribution

Wet montane forests of the Montañas del Mico (260 to 962 m elevation) and Sierra de Caral (800 to 1000 m elevation), Departamento de Izabal, Guatemala; cloud forest on the western slopes of Montaña del Cerro Azul, Sierra del Espíritu Santo, 900–1200 m elevation, Departamento de Copán, Honduras.

Geographic Occurrence

Natural Resident: Guatemala, Honduras

Comment

In the Eleutherodactylus spatulatus subgroup of the Eleutherodactylus alfredi group according to the original publication. McCranie and Smith, 2006, Herpetologica, 62: 190, suggested that this species (as Craugastor nefrens) will be found in northwestern Honduras. McCranie, 2007, Herpetol. Rev., 38: 37, summarized the departmental distribution (as Eleutherodactylus cyanochthebius) in Honduras. In the Craugastor (Hylactophryne) bocourti species series of Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 45. Padial, Grant, and Frost, 2014, Zootaxa, 3825: 123, rejected species groups within the subgenus Hylactophryne. Köhler, 2011, Amph. Cent. Am.: 140–191, provided a key to the genera and species of Brachycephaloidea (= Craugastoridae, Eleutherodactylidae) in Central America and provided maps and photographs of the species, including this one.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.