Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Craugastor emleni (Dunn, 1932)

Class: Amphibia > Order: Anura > Superfamily: Brachycephaloidea > Family: Craugastoridae > Genus: Craugastor > Species: Craugastor emleni

Eleutherodactylus ranoides nubicola Dunn and Emlen, 1932, Proc. Acad. Nat. Sci. Philadelphia, 84: 23. Holotype: ANSP 19876, by original designation. Type locality: "San Juancito, in the cloud forest at 6600 feet and . . . at 6000 feet", Departamento de Francisco Morazon, Honduras. Junior secondary homonym of Eleutherodactylus nubicola Dunn, 1926.

Eleutherodactylus ranoides emleni Dunn, 1932, Copeia, 1932: 97. Replacement name for Eleutherodactylus ranoides nubicola.

Eleutherodactylus ranoides emmetti Gorham, 1966, Das Tierreich, 85: 96. Replacement name for Eleutherodactylus ranoides nubicola.

Eleutherodactylus emleni — Campbell and Savage, 2000, Herpetol. Monogr., 14: 261.

Craugastor emleni — Crawford and Smith, 2005, Mol. Phylogenet. Evol., 35: 551, by implication; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 360.

Craugastor (Craugastor) emleni — Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 41.

Common Names

None noted.

Distribution

Elevations of 800–2000 m in the Río Choluteca drainage system of the Pacific versant of south-central Honduras between Tegucigalpa and about El Zamorano, Francisco Morazán Province.

Geographic Occurrence

Natural Resident: Honduras

Endemic: Honduras

Comment

Removed from the synonymy of Eleutherodactylus rugulosus by Campbell and Savage, 2000, Herpetol. Monogr., 14: 261, where it had been placed by Lynch, 1965, Herpetologica, 21: 102–113, and Meyer and Wilson, 1971, Contrib. Sci. Nat. Hist. Mus. Los Angeles Co., 218: 1–47. Savage, 2002, Amph. Rept. Costa Rica: 226, included this species in his Eleutherodactylus fitzingeri series, Eleutherodactylus rugulosus group. See account by McCranie and Wilson, 2002, Amph. Honduras: 382–386. McCranie, 2007, Herpetol. Rev., 38: 37, summarized the departmental distribution in Honduras. In the Craugastor punctariolus species series of Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 42, and of Padial, Grant, and Frost, 2014, Zootaxa, 3825: 123. See map, description of geographic range and habitat, and conservation status (as Eleutherodactylus emleni) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 335, who noted that this species may be extinct. Köhler, 2011, Amph. Cent. Am.: 140–191, provided a key to the genera and species of Brachycephaloidea (= Craugastoridae, Eleutherodactylidae) in Central America and provided maps and photographs of the species, including this one.

External links:

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.