Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Craugastor gulosus (Cope, 1875)

Class: Amphibia > Order: Anura > Superfamily: Brachycephaloidea > Family: Craugastoridae > Genus: Craugastor > Species: Craugastor gulosus

Lithodytes gulosus Cope, 1875 "1876", J. Acad. Nat. Sci. Philadelphia, Ser. 2, 8: 112. Holotype: USNM 32590 according to Cochran, 1961, Bull. U.S. Natl. Mus., 220: 65. Type locality: "on a spur of the Pico Blanco, at 6000 feet elevation", Costa Rica. Corrected to "Cerro Utyum, 6000 ft, Cantón de Talamanca, Provincia de Limón; 1829 m", Costa Rica, by Savage, 1974, Rev. Biol. Tropical, 22: 110 and by Savage and Myers, 2002, Am. Mus. Novit., 3357: 25. See discussion of type locality by Arias and Chaves, 2014, Mesoam. Herpetol., 1: 176–180, who concluded that the type locality is actually ""Near an unnamed hill close to the Río Lari canyon, elev. 1,830 m", Provincia de Limón, Costa Rica.

Hylodes gulosus — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 211; Brocchi, 1882, Miss. Scient. Mex. Amer. Centr., Rech. Zool., 3(2, livr. 2): 57.

Eleutherodactylus gulosus — Stejneger, 1904, Annu. Rep. U.S. Natl. Mus. for 1902: 582–583, by implication; Savage and Myers, 2002, Am. Mus. Novit., 3357: 10.

Craugastor gulosus — Crawford and Smith, 2005, Mol. Phylogenet. Evol., 35: 551, by implication; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 360.

Craugastor (Craugastor) gulosus — Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 38.

Common Names

None noted.

Distribution

Humid premontane and possibly lower montane areas of the southern Cordillera de Talamanca of Costa Rica and Panama, 1000–1873 m elevation.

Geographic Occurrence

Natural Resident: Costa Rica, Panama

Comment

Removed from the synonymy of Eleutherodactylus biporcatus by Savage and Myers, 2002, Am. Mus. Novit., 3357: 10, where it had been placed by Lynch, 1975, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 38: 1-46. (it had previously been placed in the synonymy of Hylodes rugosus Peters, 1874, by Noble, 1918, Bull. Am. Mus. Nat. Hist., 38: 329. In the Eleutherodactylus biporcatus group according to Savage and Myers, 2002, Am. Mus. Novit., 3357: 1-21, who provided an account. See account by Savage, 2002, Amph. Rept. Costa Rica: 253-254, who included this species in his Eleutherodactylus fitzingeri series, Eleutherodactylus biporcatus group. In the Craugastor gulosus species series of Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 38. In the Craugastor (Craugastor) punctariolus species series of Padial, Grant, and Frost, 2014, Zootaxa, 3825: 123. See map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 304. Köhler, 2011, Amph. Cent. Am.: 140–191, provided a key to the genera and species of Brachycephaloidea (= Craugastoridae, Eleutherodactylidae) in Central America and provided maps and photographs of the species, including this one.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.