Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Craugastor loki (Shannon and Werler, 1955)

Class: Amphibia > Order: Anura > Superfamily: Brachycephaloidea > Family: Craugastoridae > Genus: Craugastor > Species: Craugastor loki

Eleutherodactylus loki Shannon and Werler, 1955, Trans. Kansas Acad. Sci., 58: 370. Holotype: Formerly F.A. Shannon 4748, now UIMNH 67057 according to Streicher, García-Vázquez, Ponce-Campos, Flores-Villela, Campbell, and Smith, 2014, Syst. Biodiversity, 12: 11. Type locality: "at an altitude of 3500 feet on Volcán San Martín", Veracruz, Mexico. Synonymy by Duellman, 1961, Univ. Kansas Publ. Mus. Nat. Hist., 15: 18-23; Köhler, Veselý, and Greenbaum, 2005 "2006", Amph. Rept. El Salvador: 53–54; but see Lynch, 2000, Rev. Acad. Colomb. Cienc. Exact. Fis. Nat., 24: 130, who considered it distinct.

Eleutherodactylus sanmartinensis Shannon and Werler, 1955, Trans. Kansas Acad. Sci., 58: 375. Holotype: formerly F.A. Shannon 4750 (now presumably in UIMNH). Type locality: "at an altitude of 3500 ft., near a water hole on Volcán San Martín, Veracruz", Mexico. Synonymy with Eleutherodactylus rhodopis by Lynch, 2000, Rev. Acad. Colomb. Cienc. Exact. Fis. Nat., 24: 130; with Eleutherodactylus loki by Lynch, 2000, Rev. Acad. Colomb. Cienc. Exact. Fis. Nat., 24: 142.

Craugastor loki — Crawford and Smith, 2005, Mol. Phylogenet. Evol., 35: 551, by implication; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 360.

Craugastor (Craugastor) loki — Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 41.

Common Names

Common Leaf-litter Frog (Campbell, 1998, Amph. Rept. N. Guatemala Yucatan Belize: 55; Streicher, García-Vázquez, Ponce-Campos, Flores-Villela, Campbell, and Smith, 2014, Syst. Biodiversity, 12: 11).

Volcan San Martin Robber Frog (Liner and Casas-Andreu, 2008, Herpetol. Circ., 38: 10).

Distribution

Between 9 and 2100 m elevation from San Luis Potosi and Veracruz along the Caribbean lowlands of Mexico to Belize and northwestern Honduras; crossing the Isthmus of Tehuantepec and thence southeast along the Pacific versant of Chiapas and Guatemala to El Salvador.

Geographic Occurrence

Natural Resident: Belize, El Salvador, Guatemala, Honduras, Mexico

Comment

See account by McCranie and Wilson, 2002, Amph. Honduras: 406–407, and In the Eleutherodactylus (Craugastor) rhodopis group according to Lynch, 2000, Rev. Acad. Colomb. Cienc. Exact. Fis. Nat., 24: 138-143. See accounts by Lee, 1996, Amph. Rept. Yucatan Peninsula: 64-65, Campbell, 1998, Amph. Rept. N. Guatemala Yucatan Belize: 55; and Lee, 2000, Field Guide Amph. Rept. Maya World: 70–71. In the Craugastor rhodopis species series, Craugastor rhodopis species group of Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 41, and of Padial, Grant, and Frost, 2014, Zootaxa, 3825: 123. Systematics reviewed by Streicher, García-Vázquez, Ponce-Campos, Flores-Villela, Campbell, and Smith, 2014, Syst. Biodiversity, 12: 1–22. Köhler, 2011, Amph. Cent. Am.: 140–191, provided a key to the genera and species of Brachycephaloidea (= Craugastoridae, Eleutherodactylidae) in Central America and provided maps and photographs of the species, including this one. See comment by Solis, Wilson, and Townsend, 2014, Mesoam. Herpetol., 1: 125.

External links:

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.