Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Craugastor rugosus (Peters, 1873)

Class: Amphibia > Order: Anura > Superfamily: Brachycephaloidea > Family: Craugastoridae > Genus: Craugastor > Species: Craugastor rugosus

Hylodes rugosus Peters, 1873, Monatsber. Preuss. Akad. Wiss. Berlin, 1873: 610. Holotype: ZMB 7812 according to Bauer, Günther, and Klipfel, 1995, in Bauer et al. (eds.), Herpetol. Contr. W.C.H. Peters: 46. Type locality: "Chiriqui", Panama (see comment under Dendrobates maculatus).

Lithodytes pelviculus Cope, 1877, Proc. Am. Philos. Soc., 17: 89. Holotype: USNM 32326 according to Cochran, 1961, Bull. U.S. Natl. Mus., 220: 66. Type locality: "said to have been found on the west coast of Central America, without more specific locality". Synonymy with Hylodes rugosus Peters, 1874, by Noble, 1918, Bull. Am. Mus. Nat. Hist., 38: 329; synonymy with Strabomantis biporcatus Peters, 1863, by Lynch, 1975, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 38: 1-46.

Hylodes pelviculus — Brocchi, 1881, Miss. Scient. Mex. Amer. Centr., Rech. Zool., 3(2, livr. 1): 55; Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 210.

Lithodytes florulentus Cope, 1893, Proc. Am. Philos. Soc., 31: 336. Syntypes: "No. 327 [G.K Cherrie collection]"; now presumed lost according to Savage and Myers, 2002, Am. Mus. Novit., 3357: 38. Type locality: "Boruca", Cantón de Buenos Aires, Provincia de Puntarenas, Costa Rica. Type locality discussed by Savage, 1974, Rev. Biol. Tropical, 22: 80. Synonymy with Eleutherodactylus biporcatus by Lynch, 1975, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 38: 1-46. Synonymy with Hylodes rugosus Peters, 1873, by Savage and Myers, 2002, Am. Mus. Novit., 3357: 38.

Hylodes florulentus — Günther, 1900, Biol. Centr. Amer., Rept. Batr., Vol. 7, Part 157: 234.

Eleutherodactylus pelviculus — Stejneger, 1904, Annu. Rep. U.S. Natl. Mus. for 1902: 582-583, by implication.

Eleutherodactylus rugosus — Stejneger, 1904, Annu. Rep. U.S. Natl. Mus. for 1902: 582-583, by implication; Noble, 1918, Bull. Am. Mus. Nat. Hist., 38: 339.

Eleutherodactylus florulentus — Stejneger, 1904, Annu. Rep. U.S. Natl. Mus. for 1902: 582-583, by implication; Taylor, 1952, Univ. Kansas Sci. Bull., 35: 765.

Eleutherodactylus (Craugastor) florulentus — Hedges, 1989, in Woods (ed.), Biogeograph. W. Indies: 317, by implication.

Craugastor florulentus — Crawford and Smith, 2005, Mol. Phylogenet. Evol., 35: 551, by implication; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 360.

Craugastor (Craugastor) rugosus — Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 38.

Common Names

Veragua Robber Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 73).

Distribution

Lowlands and premontane slopes of the Pacific versant from the Rio Carara, Puntarenas Province, Costa Rica (9° 47′ N) to extreme western Panama (10–1220 m).

Geographic Occurrence

Natural Resident: Costa Rica, Panama

Comment

Removed from the synonymy of Eleutherodactylus biporcatus by Savage and Myers, 2002, Am. Mus. Novit., 3357: 38 (and who provided an account on page 38-39), where it had been placed by Lynch, 1975, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 38: 1-46. See account by Savage, 2002, Amph. Rept. Costa Rica: 255-256, who included this species in his Eleutherodactylus fitzingeri series, Eleutherodactylus biporcatus group. In the Craugastor gulosus species series of Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 38. In the Craugastor (Craugastor) punctariolus species series of Padial, Grant, and Frost, 2014, Zootaxa, 3825: 123. Köhler, 2011, Amph. Cent. Am.: 140–191, provided a key to the genera and species of Brachycephaloidea (= Craugastoridae, Eleutherodactylidae) in Central America and provided maps and photographs of the species, including this one.

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.