Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Pristimantis quaquaversus (Lynch, 1974)

Class: Amphibia > Order: Anura > Superfamily: Brachycephaloidea > Family: Craugastoridae > Subfamily: Pristimantinae > Genus: Pristimantis > Species: Pristimantis quaquaversus

Eleutherodactylus quaquaversus Lynch, 1974, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 31: 9. Holotype: KU 123745, by original designation. Type locality: "south slope of the Cordillera del Dué above the Río Coca, Provincia Napo, Ecuador, 1150 m."

Eleutherodactylus (Eleutherodactylus) quaquaversus — Lynch, 1996, in Powell and Henderson (eds.), Contr. W. Indian Herpetol.: 154; Lynch and Duellman, 1997, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 23: 231.

Pristimantis quaquaversus — Heinicke, Duellman, and Hedges, 2007, Proc. Natl. Acad. Sci. USA, Suppl. Inform., 104: Table 2.

Pristimantis (Pristimantis) quaquaversus — Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 121.

Common Names

Variable Rainfrog (Coloma and Duellman, 2025, Amph. Ecuador. Vol. 3: xxxvi).

Cutín Variable (Spanish: Coloma and Duellman, 2025, Amph. Ecuador. Vol. 3: xxxvi).

Rio Coca Robber Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 78).

Distribution

Lower Amazonian slopes (200 to 2362 m) of the Andes from northern Ecuador south to the Cordillera de Cutucú and Cordillera del Condor, and adjacent Peru; likely to extend into adjacent Colombia.

Geographic Occurrence

Natural Resident: Ecuador, Peru

Likely/Controversially Present: Colombia

Comment

In the Eleutherodactylus (Eleutherodactylus) martinicensis series, Eleutherodactylus unistrigatus group according to Lynch and Duellman, 1997, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 23: 231. Duellman, 1978, Misc. Publ. Mus. Nat. Hist. Univ. Kansas, 65: 100–101, provided a brief account and characterization of the call. Lynch and Duellman, 1980, Misc. Publ. Mus. Nat. Hist. Univ. Kansas, 69: 53–54, also provided a brief account. Peruvian record supplied by Duellman and Mendelson, 1995, Univ. Kansas Sci. Bull., 55: 329–376. See account by Duellman and Pramuk, 1999, Sci. Pap. Nat. Hist. Mus. Univ. Kansas, 13: 1–78. In the Pristimantis (Pristimantis) frater species group of Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 121. Not assignable to a species group according to Padial, Grant, and Frost, 2014, Zootaxa, 3825: 128. See Duellman and Lehr, 2009, Terrest.-breeding Frogs in Peru: 222–223, for brief account. Reyes-Puig, Reyes-Puig, and Yánez-Muñoz, 2013, Avanc. Cienc. Ingen., Quito, Secc. B,, 5(2): B5–B13, commented on the range in Ecuador. Brito-M., Batallas-Revelo, and Yánez-Muñoz, 2017, Neotropical Biodiversity, 3: 125–156, provided a record from Morona Santiago, Ecuador and brief natural history notes. Coloma and Duellman, 2025, Amph. Ecuador. Vol. 3: 336–340, provided an account with photographs which summarized morphological identification, systematics, natural history, distribution (including a dot map for Ecuador), and conservation.

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to Ecuador see FaunaWebEcuador: Anfibios del Ecuador