Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Pristimantis luscombei (Duellman and Mendelson, 1995)

Class: Amphibia > Order: Anura > Superfamily: Brachycephaloidea > Family: Strabomantidae > Subfamily: Pristimantinae > Genus: Pristimantis > Species: Pristimantis luscombei

Eleutherodactylus luscombei Duellman and Mendelson, 1995, Univ. Kansas Sci. Bull., 55: 354. Holotype: URP (W.E. Duellman 59957), by original designation. Type locality: "study zone at 1.5 km N Teniente Lopez, (02° 35′ 39.6″ S, 76° 06′ 55.0″ W, 312 m), Provincia Loreto, Departamento Loreto, Peru".

Eleutherodactylus (Eleutherodactylus) luscombei — Lynch and Duellman, 1997, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 23: 227.

Pristimantis luscombei — Heinicke, Duellman, and Hedges, 2007, Proc. Natl. Acad. Sci. USA, Suppl. Inform., 104: Table 2.

Pristimantis (Pristimantis) luscombei — Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 128.

Pristimantis achuar Elmer and Cannatella, 2008, Zootaxa, 1784: 13. Holotype: QCAZ25463, by original designation. Type locality: "Kapawi Jungle Lodge, Pastaza province, Ecuador (S 02° 32.32′, W 76° 51.50′, altitude 239 m)". Synonymy by Ortega-Andrade and Venegas, 2014, Zootaxa, 3895: 34–35.

English Names

None noted.

Distribution

Amazonas and Loreto, Peru, and adjacent provinces of Morona-Santiago and Pastaza in Amazonian Ecuador; also reported in Cruzeiro do Sul, Acre, Brazil, and from western Putumayo, Colombia.

Comment

In the Eleutherodactylus unistrigatus group according to the original publication. In the Eleutherodactylus (Eleutherodactylus) martinicensis series, Eleutherodactylus unistrigatus group of Lynch and Duellman, 1997, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 23: 227. In the Pristimantis (Pristimantis) unistrigatus species group of Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 128. Not assignable to a species group according to Padial, Grant, and Frost, 2014, Zootaxa, 3825: 127. See Duellman and Lehr, 2009, Terrest.-breeding Frogs in Peru: 190, for brief account. Ortega-Andrade and Valencia, 2010, Herpetol. Notes, 3: 251–256, provided the first records for Ecuador. Betancourth-Cundar and Gutiérrez-Zamora, 2010, Ecotrópicos, 23: 61–78m reported the species (as Pristimantis achuar) from the Centro Experimental Amazonico, western Putumayo, Colombia. Ortega-Andrade and Venegas, 2014, Zootaxa, 3895: 34–35, provided an account, placed Pristimantis achuar in synonymy, and named Pristimantis miktos from within the range of this species, which although it did not change the understood range of this species, certainly changed the diagnosis of the species. Pristimantis luscombei (as Pristimantis achuar) was confused with Pristimantis ockendeni prior to its naming. López-Rojas, Ramalho, Susçuarana, and Souza, 2013, Check List, 9: 1548–1551, provided a range extension (as Pristimantis achuar) into Acre, Brazil.

External links:

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For additional sources of information from other sites search Google
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist
For additional information specific to Ecuador see FaunaWebEcuador: Anfibios del Ecuador
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.