Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Nannophryne variegata Günther, 1870

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Nannophryne > Species: Nannophryne variegata

Nannophryne variegata Günther, 1870, Proc. Zool. Soc. London, 1870: 402. Syntypes: BMNH 1869.5.3.50-51, 1869.5.3.59 (reregistered , 1947.2.21.98), and BMNH 1868.9.22.3 (reregistered 1947.2.21.96), and 2 not traced, according to Hoogmoed, 1985, in Frost (ed.), Amph. Species World: 63. Type localities: "Puerto Bueno, Port Grappler, and in Eden Harbour", Magallanes, Chile; Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 293, listed at least 6 specimens as syntypes from "Chili", "Puerto Bueno", "Port Grappler", and "Eden Harbour"; restricted to "Puerto Bueno (Magallanes), Chile" by Cei, 1962, Batr. Chile: 51.

Bufo variegatus — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 293; Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 36.

Phryniscus variegatus — Boulenger, 1894, Ann. Mag. Nat. Hist., Ser. 6, 14: 374, by implication.

Bufo albigularis Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 32. Syntypes: MNHNC (numerous individuals), according to the original publication. Type locality: "Andibus provinciae Cautin" Chile. Considered a nomen dubium by Nieden, 1923, Das Tierreich, 46: 146. Synonymy by Cei, 1958, Invest. Zool. Chilen., 4: 267; Gallardo, 1962, Physis, Buenos Aires, 23: 99; Cei, 1980, Monit. Zool. Ital., N.S., Monogr., 2: 160; Lavilla, 1994 "1992", Acta Zool. Lilloana, 42: 67.

Bufo trivittatus Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 38. Syntypes: MNHNC (3 specimens), according to the original publication. Type locality: "Araucanía", Chile. Considered a nomen dubium by Nieden, 1923, Das Tierreich, 46: 146. Synonymy by Cei, 1958, Invest. Zool. Chilen., 4: 267; Gallardo, 1962, Physis, Buenos Aires, 23: 99; Cei, 1980, Monit. Zool. Ital., N.S., Monogr., 2: 160.

Nannophryne variegata — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 364.

Nannophryne variegatus — Pramuk, Robertson, Sites, and Noonan, 2008, Global Ecol. Biogeograph., 17: 76.

Common Names

Eden Harbour Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 44).

Patagonian Toad (Ananjeva, Borkin, Darevsky, and Orlov, 1988, Dict. Amph. Rept. Five Languages: 42).

Distribution

Neuquén to southern Río Negro in Argentina as well as adjacent Chile (Región del Biobío) southward to the Magellanic Islands of Chile (Península de Brunswick) and Argentina (see comment); possibly to Peru (see comment).

Geographic Occurrence

Natural Resident: Argentina, Chile, Peru

Comment

See comment under Nannophryne. In the Bufo variegatus group of Martin, 1972, in Blair (ed.), Evol. Genus Bufo: 59. Inclusion in the Bufo spinulosus group of Blair, 1972, Evol. Genus Bufo: 346, is controversial; see Pramuk and Kadivar, 2003, Herpetologica, 59: 270–271, for discussion. For discussion and synonymy see Gallardo, 1962, Physis, Buenos Aires, 23: 93–102. For account see Cei, 1962, Batr. Chile: 51-52, and Cei, 1980, Monit. Zool. Ital., N.S., Monogr., 2: 160–164. Cei, 1980, declined to place this species in any of the species groups. Biolé and Williams, 1985 "1984", Neotropica, 30: 243–244, discussed some aspects of geographic variation. The record for Peru by Rodríguez, Córdova, and Icochea, 1993, Publ. Mus. Hist. Nat. Univ. Nac. Mayor San Marcos, 45: 1–22, requires confirmation as being ecologically and geographically very isolated from other records. See Lobos, Vidal-Maldonado, Correa-Quezada, Labra-Lillo, Díaz-Páez, Charrier, Rabanal, Díaz-Vidal, and Tala, 2013, Anf. Chile Conserv. 1–104, for photograph, comments on conservation status, and range. See Charrier, 2019, Anf. Bosques Centro Sur y Patagonia de Chile: 214–217, for account (morphology, habitat, natural history) and polygon distribution map.

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