Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Pelophryne signata (Boulenger, 1895)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Pelophryne > Species: Pelophryne signata

Nectophryne signata Boulenger, 1895 "1894", Proc. Zool. Soc. London, 1894: 645. Syntypes: BMNH (2 specimens), by original designation; these being BMNH 1894.6.30.135–136 (reregistered 1947.2.19.25-26) according to museum records. Type locality: "Robong [= Rabong] Mt., Kapunas district, Dutch Borneo", Kalimantan, Borneo.

Nectophryne exigua Boettger, 1900, Abh. Senckenb. Naturforsch. Ges., 25: 394. Holotype: SMF 3737 (formerly 1280a) according to Mertens, 1967, Senckenb. Biol., 48(A): 40. Type locality: "Baramfluss, Nord-Borneo" (= Baram River, Sarawak), Malaysia (Borneo). Synonymy by Inger and Stuebing, 2009, Raffles Bull. Zool., 57: 532.

Pelophryne signata — Barbour, 1938, Proc. Biol. Soc. Washington, 51: 193; Inger, 1985, in Frost (ed.), Amph. Species World: 38; by implication; Malkmus, Manthey, Vogel, Hoffmann, and Kosuch, 2002, Amph. Rept. Mount Kinabalu: 84.

Common Names

Saint Andrew's Cross Toadlet (Lim and Lim, 1992, Guide Amph. Rept. Singapore: 24).

Short-legged Dwarf Toad (Das, Jankowski, Makmor, and Haas, 2007, Mitt. Hamburg. Zool. Mus. Inst., 104: 148; Das, 2007, Amph. Rept. Brunei: 32).

Dwarf Toad (Grismer, 2012, Field Guide Amph. Rept. Seribuat Arch.: 49).

Lowland Dwarf Toad (Haas, Das, Hertwig, Bublies, and Schulz-Schaeffer, 2022, Guide to the Tadpoles of Borneo: 255).

Distribution

Tropical lowland forests of northern Borneo (Sarawak, Malaysia; Brunei) and adjacent Kalimantan, Indonesia. Serasan and Bunguran Is., Natuna Is., Indonesia. See comment regarding East Malaysia and Singapore populations.

Geographic Occurrence

Natural Resident: Brunei, Indonesia, Malaysia, Malaysia, East (Sarawak and/or Sabah)

Comment

See Inger, 1966, Fieldiana, Zool., 52: 85 (as Pelophryne brevipes), for synonymy and distribution although this should be used with caution as population from West Malaysia is now Pelophryne ingeri. See brief account (as Pelophryne brevipes) and photo by Manthey and Grossmann, 1997, Amph. Rept. Südostasiens: 41-43, for the Borneo population. Removed from the synonym of Pelophryne brevipes by Inger, 1985, in Frost (ed.), Amph. Species World: 38, where it had been placed by Inger, 1966, Fieldiana, Zool., 52: 85. See comment under Pelophryne brevipes. Das, Jankowski, Makmor, and Haas, 2007, Mitt. Hamburg. Zool. Mus. Inst., 104: 148, provided a brief description. See statement of geographic range, habitat, and conservation status in Chan and Grismer, 2019, BMC Evol. Biol., 19(95): 3, suggested on the basis of sequence divergence between Borneo and Peninsular Malaysia that this binominal represents a species complex (see Pelophryne ingeri, which appears to resolve this–DRF). Matsui, 2019, Curr. Herpetol., Kyoto, 38: 128–139, discussed the confusing history of the application of this name to populations now known as Pelophryne ingeri and Pelophryne brevipes. Haas, Das, Hertwig, Bublies, and Schulz-Schaeffer, 2022, Guide to the Tadpoles of Borneo: 255–257, summarized the knowledge of habitat, reproduction, larval morphology and coloration. Herlambang, Riyanto, Munir, Hamidy, Kimura, Eto, and Mumpuni, 2022, Treubia, 49: 78, reported the species from Bungaran and Serasan Is., Natuna Is., Indonesia.

External links:

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For access to general information see Wikipedia
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For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.