Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Ceratophrys aurita (Raddi, 1823)

Class: Amphibia > Order: Anura > Family: Ceratophryidae > Genus: Ceratophrys > Species: Ceratophrys aurita

Bufo auritus Raddi, 1823, Mem. Mat. Fis. Soc. Ital. Sci. Modena, 19: 71. Type(s): Not stated, although presumably MZUF. Type locality: "Montagne d'Estrella, a due giornate circa da Rio-janeiro", Brazil. Given by Bokermann, 1966, Lista Anot. Local. Tipo Anf. Brasil.: 18, as "Serra da Estrela, Inhomirin, Rio de Janeiro", Brazil.

Rana macrocephala Wied-Neuwied, 1824, Isis von Oken, 14: 671. Types: Not designated. Type locality: "Brasiliens". Given as "Lago da Arara on the lower Rio Mucuri" Brazil, by Vanzolini and Myers, 2015, Bull. Am. Mus. Nat. Hist., 395: 77. Synonymy by W.R. Heyer (personal commun.) See comments by Vanzolini and Myers, 2015, Bull. Am. Mus. Nat. Hist., 395: 76.

Ceratophrys varius Wied-Neuwied, 1824, Isis von Oken, 14: 673. Types: Specimens not designated but including specimens figured by Seba, 1734, Locuplet. Rer. Nat. Thesaur. Descript. Icon. Exp. Univ. Phys. Hist., 1: Pl. 72, figs. 1-2. Type locality: "Brasilien"; restricted to "Itabapuana, Rio de Janeiro", Brazil by Bokermann, 1966, Lista Anot. Local. Tipo Anf. Brasil.: 26. Synonymy by Bokermann, 1965, Neotropica, 11: 12.

Ceratophrys dorsata Wied-Neuwied, 1824, Isis von Oken, 14: 673. Types: Not designated. Type locality: Not stated beyond obviously being between Bahia and Rio de Janeiro, Brazil; given as "aus der Urwäldern am Itabapuana-Fluss im Staate Rio de Janeiro", by Müller, 1927, Abh. Senckenb. Naturforsch. Ges., 40: 268. See Bokermann, 1966, Lista Anot. Local. Tipo Anf. Brasil.: 25, for comments on type locality. Synonymy (with Ceratophrys varia) by Cochran, 1955 "1954", Bull. U.S. Natl. Mus., 206: 232-233, and Cei, 1956, Invest. Zool. Chilen., 3: 41. Synonymy by Miranda-Ribeiro, 1920, Rev. Mus. Paulista, São Paulo, 12: 297; Bokermann, 1965, Neotropica, 11: 9-12.

Ceratophrys dorsata — Wied-Neuwied, 1827, Abbild. Naturgesch. Brasil., Heft 10: pl. 45, fig. 1; Schinz, 1833, Naturgesch. Abbild Rept.: 226; Tschudi, 1838, Classif. Batr.: 81; S

Stombus dorsatus — Gravenhorst, 1829, Delic. Mus. Zool. Vratislav., 1: 49.

Ceratophris clipeata Cuvier, 1829, Regne Animal., Ed. 2, 2: 106. Types: Not designated, presumably deposited in MNHNP, but not recorded there now. Type locality: Not stated; given as Amerique meridionale by Gorham, 1966, Das Tierreich, 85: 39. Nomen nudum. Synonymy (with Ceratophrys dorsata) by Duméril and Bibron, 1841, Erp. Gen., 6: 430; Cei, 1956, Invest. Zool. Chilen., 3: 41.

Ceratophrys clypeata — Cocteau, 1835, Mag. Zool., Guérin, 5: pl. 8, fig. 2.

Ceratophrys varia — Cocteau, 1835, Mag. Zool., Guérin, 5: pl. 8, fig. 2.

Ceratophryne dorsata — Schlegel, 1858, Handl. Dierkd., 2: 57.

Ceratophrys aurita — Bokermann, 1966, Lista Anot. Local. Tipo Anf. Brasil.: 18.

Ceratophrys (Ceratophrys) aurita — Lynch, 1982, Syst. Zool., 31: 166-179.

Common Names

Wied's Frog (Ceratophrys varia [no longer recognized]: Cochran, 1961, Living Amph. World: 63).

Wied's Frog (Ananjeva, Borkin, Darevsky, and Orlov, 1988, Dict. Amph. Rept. Five Languages: 78).

Brazilian Horned Frog (Ananjeva, Borkin, Darevsky, and Orlov, 1988, Dict. Amph. Rept. Five Languages: 78; Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 71).

Distribution

Minas Gerais and Bahia to Rio Grande do Sul, Brazil.

Geographic Occurrence

Natural Resident: Brazil

Endemic: Brazil

Comment

See account (as Ceratophrys varia) by Cochran, 1955 "1954", Bull. U.S. Natl. Mus., 206: 232-234, and (as Ceratophrys aurita) by Heyer, Rand, Cruz, Peixoto, and Nelson, 1990, Arq. Zool., São Paulo, 31: 289. Izecksohn and Carvalho-e-Silva, 2001, Anf. Municipio Rio de Janeiro: 69, provided a brief account and photo. Faivovich, Nicoli, Blotto, Pereyra, Baldo, Barrionuevo, Fabrezi, Wild, and Haddad, 2014, S. Am. J. Herpetol., 9: 207–227, suggested that Ceratophrys aurita is the sister taxon of Ceratophrys joazeirensis. Forti, Haddad, Leite, Drummond, Assis, Crivellari, Mello, Garcia, Zornosa-Torres, and Toledo, 2019, PeerJ, 7(e7612): 1–39, reported on advertisement call. Mira-Mendes, Dias, Costa, Costa, Silva, Weber, and Zina, 2022, Zootaxa, 5124: 594–600, reported on the external and internal oral cavity anatomy of the larva.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.