Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Ceratophrys joazeirensis Mercadal de Barrio, 1986

Class: Amphibia > Order: Anura > Family: Ceratophryidae > Genus: Ceratophrys > Species: Ceratophrys joazeirensis

Ceratophrys joazeirensis Mercadal de Barrio, 1986, Amphibia-Reptilia, 7: 320. Holotype: NHMW 4582, by original designation. Type locality: "Joazeiro (Estado de Bahía), NE de Brasil".

Common Names

Joazeiro Horned Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 71).

Distribution

Brazilian states of Minas Gerais (Fazenda Jatobá, Buritizeiro), Bahia (Joazeiro), Paraiba (Municipality of Araruna), Rio Grande do Norte (Santa Maria Municipality), Pernambuco (Municipality of Triunfo), Alagoas, and Sergipe.

Geographic Occurrence

Natural Resident: Brazil

Endemic: Brazil

Comment

An octoploid species most closely related to Ceratophrys ornata according to original publication. In the subgenus Ceratophrys by implication. Vieira, Arzabe, and Vieira, 2006, Check List, 2(2): 28–29, provided records from Paraíba, Brazil. Santos, Silva, Campos, and Quirino, 2009, Herpetol. Rev., 40: 107, provided the first record from Pernambuco, Brazil. Vieira, Silva, and Arzabe, 2006, Zootaxa, 1320: 57–68, discussed cranial morphology and karyology in light of relationships to other Ceratophrys. Jorge, Kokubum, Sales, and Freire, 2012, Herpetol. Rev., 43: 297–298, provided a record for Rio Grande do Norte, Brazil. Zaidan and Leite, 2012, Zootaxa, 3540: 65–66, reported on the advertisement call. Maciel, Kokubum, Braga, Queiróz, and Matsushita, 2013, Herpetol. Notes, 6: 447-450, provided a dot map, a new record for Minas Gerais, and described the advertisement call. Faivovich, Nicoli, Blotto, Pereyra, Baldo, Barrionuevo, Fabrezi, Wild, and Haddad, 2014, S. Am. J. Herpetol., 9: 207–227, suggested that this species is the sister taxon of Ceratophrys aurita. Jorge, Sales, Kokubum, and Freire, 2015, Phyllomedusa, 14: 147–156, reported on natural history and discussed the range. Almeida, Nascimento, Torquato da Silva, Lisboa, Tiburcio, Palmeira, Lima, and Mott, 2016, Herpetol. Notes, 9: 134, provided a record for Alagoas, Brazil. Dubeux, Silva, Nascimento, Gonçalves, and Mott, 2019, Rev. Nordestina Zool., 12: 18–52, noted that the larva remains undescribed.

External links:

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For access to general information see Wikipedia
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For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.