Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Cycloramphus heyeri Segalla, Berneck, Canedo, Caramaschi, Cruz, Garcia, Grant, Haddad, Lourenço, Mângia, Mott, Nascimento, Toledo, Werneck, and Langone, 2021

Class: Amphibia > Order: Anura > Family: Cycloramphidae > Genus: Cycloramphus > Species: Cycloramphus heyeri

Cycloramphus carvalhoi Heyer, 1983, Arq. Zool., São Paulo, 30: 297. Holotype: L 132, by original designation; this corrected to MNRJ 33852 by Caramaschi and Pombal, 2023, Zootaxa, 5339: 469. Type locality: "Brasil; Rio de Janeiro, Brejo da Lapa, Alto Itatiaia, 2200 m". Augmented by Caramaschi and Pombal, 2023, Zootaxa, 5339: 469, to "Brejo da Lapa (22°21’28”S, 44°44’10”W; Datum WGS 84; 2,150 m a.s.l.), Parque Nacional do Itatiaia, Municipality of Itamonte, State of Minas Gerais, Brasil". Date of receipt October 1983 according to Segalla, Berneck, Canedo, Caramaschi, Cruz, Garcia, Grant, Haddad, Lourenço, Mângia, Mott, Nascimento, Toledo, Werneck, and Langone, 2021, Herpetol. Brasil., 10: 124, who noted this as a junior subjective homonym of Cycloramphus carvalhoi Izecksohn 1983.

Cycloramphus heyeri Segalla, Berneck, Canedo, Caramaschi, Cruz, Garcia, Grant, Haddad, Lourenço, Mângia, Mott, Nascimento, Toledo, Werneck, and Langone, 2021, Herpetol. Brasil., 10: 124. Replacement name for Cycloramphus carvalhoi Heyer, 1983, a junior subjective homonym of Zachaenus carvalhoi Izecksohn, 1983, when both are in Cycloramphus. See summary comment by Caramaschi and Pombal, 2023, Zootaxa, 5339: 469.

English Names

Carvalho's Button Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 71).

Distribution

Known from the type locality (Alto Itatiaia, Rio de Janeiro) and about 200 km to the southwest in the Municipality of Campos do Jordão (São Paulo), Brazil.

Geographic Occurrence

Natural Resident: Brazil

Endemic: Brazil

Comment

Martins, Gomes, Bernardo, and Suarez, 2007, Herpetol. Rev., 38: 473-474, provided the record from the state of São Paulo, Brazil (as Cycloramphus carvalhoi).

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For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.