Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Rheobates palmatus (Werner, 1899)

Class: Amphibia > Order: Anura > Superfamily: Dendrobatoidea > Family: Aromobatidae > Subfamily: Anomaloglossinae > Genus: Rheobates > Species: Rheobates palmatus

Phyllobates (Hypodictyon) palmatus Werner, 1899, Verh. Zool. Bot. Ges. Wien, 49: 479. Holotype: Originally ZIUG; now ZFMK 28134, according to Böhme and Bischoff, 1984, Bonn. Zool. Monogr., 19: 180. Type locality: "Fusagasugá, 1700 m, (bei Bogotá)", Departamento Cundinamarca, Colombia.

Hylixalus granuliventris Boulenger, 1919, Proc. Zool. Soc. London, 1919: 79–81. Holotype: BMNH 1947.2.14.28 (originally 1919.3.6.32), by museum records. Type locality: "Bogotá", Cundinamarca, Colombia. Synonymy by Dunn, 1957, Copeia, 1957: 77; Cochran and Goin, 1970, Bull. U.S. Natl. Mus., 288: 52.

Syrrhophus palmatus — Barbour and Noble, 1920, Bull. Mus. Comp. Zool., 63: 401.

Hyloxalus granuliventris — Hellmich, 1939, Wochenschr. Aquar. Terrarienkd., Braunschweig, 36: 536; Hellmich, 1940, Zool. Anz., 131: 114; Dunn, 1944, Caldasia, 2: 403.

Hyloxalus palmatus — Dunn, 1944, Caldasia, 2: 403.

Prostherapis palmatus — Dunn, 1957, Copeia, 1957: 77.

Phyllobates palmatus — Cochran, 1966, Mem. Inst. Butantan, São Paulo, 33: 64.

Prostherapis granuliventris —Dunn In Savage, 1968, Copeia, 1968: 6, by implication.

Colostethus granuliventris — Edwards, 1971, Proc. Biol. Soc. Washington, 84: 148.

Colostethus palmatus — Edwards, 1971, Proc. Biol. Soc. Washington, 84: 148.

Colostethus palmatus granuliventris — Gorham, 1974, Checklist World Amph.: 115.

Colostethus palmatus palmatus — Gorham, 1974, Checklist World Amph.: 115.

Rheobates palmatus — Grant, Frost, Caldwell, Gagliardo, Haddad, Kok, Means, Noonan, Schargel, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 299: 159.

Common Names

Palm Rocket Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 49).

Distribution

Cordillera Oriental (eastern slope in Boyacá, Cundinamarca, and Meta; western slope in Norte de Santander south to Huila; eastern slope of the Cordillera Central (Caldas, Tolima, and Huila), northern part of the Cordillera Occidental and Cordillera Central (southern Córdoba, Sucre, Bolívar, and northern Antioquia), and Serranía de La Macarena at elevations of 250–2520 m elevation, Colombia.

Geographic Occurrence

Natural Resident: Colombia

Endemic: Colombia

Comment

Prior to the revision of Grant, Frost, Caldwell, Gagliardo, Haddad, Kok, Means, Noonan, Schargel, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 299, in the Colostethus fuliginosus group (Group VI) of Rivero, 1990 "1988", Mem. Soc. Cienc. Nat. La Salle, 48: 3–32. Lynch and Renjifo, 2001, Guia Anf. Rept. Bogota: 43, presented a brief account and photograph. Bernal, Guarnizo, and Lüddecke: 395–408, reported on geographic variation in advertisement call. Lynch, 2006, Caldasia, 28: 139, characterized the range in Colombia. Sánchez, 2013, Zootaxa, 3637: 569–591, provided information on larval morphology. Muñoz-Ortiz, Velásquez-Álvarez, Guarnizo, and Crawford, 2015, J. Biogeograph., 42: 193–205. Guarnizo, Paz, Muñoz-Ortiz, Flechas-Hernández, Méndez-Narváez, and Crawford, 2016, PLoS One, 10(5: e0127312): 1–20, suggested on the basis of molecular evidence that three species likely exist in Colombia under this binomial.Grant, Rada, Anganoy-Criollo, Batista, Dias, Jeckel, Machado, and Rueda-Almonacid, 2017, S. Am. J. Herpetol., 12 (Special Issue): 35, provided a tree which places Rheobates pseudopalmata within multiple lineages of nominal Rheobates palmatus, suggesting that the latter is a species complex. Jerez and Yara-Contreras, 2018, Catal. Anf. Rept. Colombia, Medellín, 4(1): 68–78, provided a detailed account. Ovalle-Pacheco, Camacho-Rozo, and Arroyo, 2019, Check List, 15: 390, extended the elevational range. Genty, Guarnizo, Ramírez, Barrientos, and Crawford, 2022, J. Zool. Syst. Evol. Res., 2022 (6774225): 1–20, discussed cryptic species under this name.

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For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.