Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Ameerega pepperi Brown and Twomey, 2009

Class: Amphibia > Order: Anura > Superfamily: Dendrobatoidea > Family: Dendrobatidae > Subfamily: Colostethinae > Genus: Ameerega > Species: Ameerega pepperi

Ameerega pepperi Brown and Twomey, 2009, Zootaxa, 2049: 16. Holotype: MUSM 2640, by original designation. Type locality: "Provincia Tocache, Departamento San Martín, Peru, 2 km NE of San Francisco, 980 m elevation, 8° 18′ 30.3″ S, 76° 40′ 37.6″ W. Found on the ground near a small waterfall."

Common Names

Pepper's Poison Frog (Geralds, Twomey, Deutsch, Chávez, and Brown, 2026, Zootaxa, 5759: 311).

Distribution

Premontane elevations (407–825 m) throughout the upper Huallaga Valley from south of Río Huayabamba near Huicungo to the southern San Martín border; most records lie along the road between Tingo María and Juanjui on the river’s eastern bank (western slope of Cordillera Azul), but populations also exist on the western bank (eastern Andean slope) at San Francisco and Huicungo.

Geographic Occurrence

Natural Resident: Peru

Endemic: Peru

Comment

The advertisement call was described in the original publication as well as external morphology and natural history. In the Ameerega bassleri species group of Guillory, French, Twomey, Chávez, Prates, von May, De la Riva, Lötters, Reichle, Serrano-Rojas, Whitworth, and Brown, 2020, Mol. Phylogenet. Evol., 142 (106638): 1–13. Geralds, Twomey, Deutsch, Chávez, and Brown, 2026, Zootaxa, 5759: 295–320, revised the Ameerega bassleri group, including a review of adult and larval morphology, systematics, phylogenetics, distribution (including a dot map of the range), and advertisement calls.

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For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observations see iNaturalist