Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Ranitomeya hwata Twomey, Melo-Sampaio, Brown, Castroviejo-Fisher, Gagliardi-Urrutia, Padial, Gutiérrez, and Chaparro, 2025

Class: Amphibia > Order: Anura > Superfamily: Dendrobatoidea > Family: Dendrobatidae > Subfamily: Dendrobatinae > Genus: Ranitomeya > Species: Ranitomeya hwata

Ranitomeya hwata Twomey, Melo-Sampaio, Brown, Castroviejo-Fisher, Gagliardi-Urrutia, Padial, Gutiérrez, and Chaparro Auza, 2025, Zootaxa, 5701: 433. Holotype: MNRJ 91674, by original designation. Type locality: "Fazenda Experimental Catuaba, Senador Guiomard, state of Acre, Brazil (10.0727°S, 67.6239°W), 200 m elevation." ZooBank Publication registration: 8DEB156C-640C-43FF-9314-4F2E7FFF3A73

Common Names

None noted.

Distribution

Distributed throughout much of the state of Acre, Brazil, and extends into southeastern Peru in Ucayali department near the village of Sepahua; known from one site in Amazonas, Brazil (Reserva Extrativista Arapixi), Pando department in northern Bolivia, and from Rio Acre (which forms the border of Acre and Madre de Dios), making its presence in Madre de Dios department of Peru a virtual certainty. Occurs at elevations ranging from 190 to 356 m.

Geographic Occurrence

Natural Resident: Bolivia, Brazil, Peru

Comment

In the Ranitomeya vanzolinii group according to the original publication, where comparative adult and larval morphology, advertisement call, molecular markers, a dot map of its distribution, as well as sorting earlier literature under different taxonomic names. Most recently this species was considered to be outlying eastern populations of Ranitomeya sirensis. Recorded in Bolivia (as Dendrobates biolat) by De la Riva, Köhler, Lötters, and Reichle, 2000, Rev. Esp. Herpetol., 14: 57, Köhler, 2000, Bonn. Zool. Monogr., 48: 69; and Maldonado-M. and Reichle, 2007, Kempffiana, 3: 13-17. Melo-Sampaio and Souza, 2009, Herpetol. Rev., 40: 447, provided the first record (as Ranitomeya biolat) for Acre, Brazil. França and Venâncio, 2010, Biotemas, 23: 71–84, provided a record (as Ranitomeya biolat) for the municipality of Boca do Acre, Amazonas, with a brief discussion of the range. Crnobrna, Santa-Cruz Farfan, Gallegos, López-Rojas, Llanqui, Panduro Pisco, and Kelsen Arbaiza, 2023, Check List, 19: 441, provided a record (as Ranitomeya sirensis) from Ucayali Department, central-eastern Peru.

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