Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Ranitomeya imitator (Schulte, 1986)

Class: Amphibia > Order: Anura > Superfamily: Dendrobatoidea > Family: Dendrobatidae > Subfamily: Dendrobatinae > Genus: Ranitomeya > Species: Ranitomeya imitator

Dendrobates imitator Schulte, 1986, Sauria, Berlin, 8: 11. Holotype: MUSM (formerly MHNJP) 10501, by original designation. Type locality: "km 33, Carretera Tarapoto--Yurimaguas, Departamento San Martín, Peru. 550 m leg."

Ranitomeya imitator — Bauer, 1988, Het Paludarium, Netherlands, November: 6.

Dendrobates imitator yurimaguensis-imitator Schulte, 1999, Pfeilgiftfrösche: 94. Nomen nudum. See Lötters and Vences, 2001 "2000", Salamandra, 36: 247-260.

Dendrobates imitator imitator — Schulte, 1999, Pfeilgiftfrösche: 94, by implication.

Dendrobates imitator yurimaguensis Schulte, 1999, Pfeilgiftfrösche: 104. Holotype: R. Schulte Collection BD 40, by original designation; presumably destined for MUSM. Type locality: "Tieflandurwald des Río Shanusi-Paranapura Refugiums, Alto Amazonas (Peru). Ca. 180-300 m NN". Distinctiveness from Dendrobates imitator imitator rejected by Lötters and Vences, 2001 "2000", Salamandra, 36: 247-260, and Lötters, Reichle, and Jungfer, 2003, J. Nat. Hist., London, 37: 1899-1911.

Dendrobates imitator intermedius Schulte, 1999, Pfeilgiftfrösche: 93. Holotype: R. Schulte Collection BD 27, by original designation; presumably destined for MUSM. Type locality: "Huallaga Canyon, Region San Martin, Peru, 200 m NN". Synonymy by Brown and Twomey IN Brown, Twomey, Amézquita, Souza, Caldwell, Lötters, von May, Melo-Sampaio, Mejía-Vargas, Pérez-Peña, Pepper, Poelman, Sanchez-Rodriguez, and Summers, 2011, Zootaxa, 3083: 69.

Ranitomeya imitator — Grant, Frost, Caldwell, Gagliardo, Haddad, Kok, Means, Noonan, Schargel, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 299: 171.

Ranitomeya intermedia — Grant, Frost, Caldwell, Gagliardo, Haddad, Kok, Means, Noonan, Schargel, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 299: 171.

Common Names

Mimic Poison Frog (Walls, 1994, Jewels of the Rainforest: 22; Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 50).

Distribution

Amazonian rainforests of Peru (Departments: Loreto and San Martín).

Geographic Occurrence

Natural Resident: Peru

Endemic: Peru

Comment

Schulte, 1999, Pfeilgiftfrösche: 88-110, provided an account. Lötters, Jungfer, Henkel, and Schmidt, 2007, Poison Frogs: 478-483, provided an account and placed this species in their Ranitomeya ventrimaculata group. In the Ranitomeya vanzolinii species group of Brown, Twomey, Amézquita, Souza, Caldwell, Lötters, von May, Melo-Sampaio, Mejía-Vargas, Pérez-Peña, Pepper, Poelman, Sanchez-Rodriguez, and Summers, 2011, Zootaxa, 3083: 61, and who provided an account on page 68-72. Twomey, Mayer, and Summers, 2015, Herpetologica, 71: 252–259, reported on intraspecific variation. Klein, Regnet, Krings, and Rödder, 2020, Bonn Zool. Bull., 69: 191–223, reported on larval development and morphology.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.