Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Fejervarya verruculosa (Roux, 1911)

Class: Amphibia > Order: Anura > Family: Dicroglossidae > Subfamily: Dicroglossinae > Genus: Fejervarya > Species: Fejervarya verruculosa

Rana tigrina var. verruculosa Roux, 1911, Zool. Jahrb., Jena, Abt. Syst., 30: 504. Type(s): SMF 6538 (formerly 1026. 4a) designated lectotype by Mertens, 1967, Senckenb. Biol., 48(A): 46. Type locality: "Wetar, Iliwaki", Lesser Sundas, Indonesia.

Rana (Rana) verruculosa — Boulenger, 1920, Rec. Indian Mus., 20: 16. Van Kampen, 1923, Amph. Indo-Austral. Arch.: 172.

Rana cancrivora verruculosa — Dunn, 1928, Am. Mus. Novit., 315: 5.

Rana verruculosa — Mertens, 1928, Senckenb. Biol., 10: 230.

Rana cancrivora verruculosa — Mertens, 1930, Abh. Senckenb. Naturforsch. Ges., 42: 213-214.

Rana (Euphlyctis) verruculosa — Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 239, by implication.

Euphlyctis verruculosa — Poynton and Broadley, 1985, Ann. Natal Mus., 27: 124, by implication.

Limnonectes (Hoplobatrachus) verruculosus — Dubois, 1987 "1986", Alytes, 5: 60.

Limnonectes (Fejervarya) verruculosus — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 315.

Fejervarya verruculosa — Dubois and Ohler, 2000, Alytes, 18: 35. Fei, Ye, Jiang, and Xie, 2002, Herpetol. Sinica, 9: 93.

Common Names

Sundas Wart Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 100).

Banda Arc Wart Frog (Tanoyo, Reilly, Doughty, Arida, and Iskandar, 2024, Rec. W. Aust. Mus., 39: 29).

Distribution

Timor, Wetar, Sumba, Alor, Pantar, Rote, and Damar islands, Lesser Sundas, Indonesia and East Timor.

Geographic Occurrence

Natural Resident: East Timor, Indonesia

Comment

Menzies, 1987, Aust. J. Zool., 35: 388-389, and Menzies, 1992, Sci. New Guinea, 18: 119, reported this species from Papua New Guinea, in error, according to J. Menzies, in litt. to Allen Allison 9 Feb 2006. See account by Boulenger, 1920, Rec. Indian Mus., 20: 16, who resurrected this species from the synonymy of Hoplobatrachus tigerinus. Dubois, 1987 "1986", Alytes, 5: 60, considered this species to be a member of his Limnonectes (Hoplobatrachus) tigerinus group, but subsequently (Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 315) moved it into Fejervarya. Kennedi, Kusrini, Mardiastuti, and Ariefiandy, 2021, Media Konservasi , 26: 217–230, reported the species on Flores I., Indonesia, but Tanoyo, Reilly, Doughty, Arida, and Iskandar, 2024, Rec. W. Aust. Mus., 39: 15–62, who reported on the molecular phylogenetics and comparative morphology from the populations found in the Lesser Sundas, Indonesia, regarded this in error for Fejervarya iskandari, and regarded that species as restricted to the Banda Arc Islands.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist