Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Limnonectes dabanus (Smith, 1922)

Class: Amphibia > Order: Anura > Family: Dicroglossidae > Subfamily: Dicroglossinae > Genus: Limnonectes > Species: Limnonectes dabanus

Rana macrognathus dabana Smith, 1922, J. Nat. Hist. Soc. Siam, 4: 221. Syntypes: BMNH (2 specimens—formerly M. Smith 4842 and 4845), by original designation. Type locality: "Daban, 200 metres altitude, Langbian plateau, S. Annam" (now in Loc Chau Commune, Bao Loc City, Lam Dong Province, Vietnam, elevation 200 m, according to Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 23).

Rana toumanoffi Bourret, 1941, Annexe Bull. Gen. Instr. Publique, Hanoi, 1941: 23. Holotype: R. Bourret 285, by original designation; now MNHNP 1948.126 according to Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 39. Type locality: "Mimot (Cambodge)". Synonymy by Stuart, Sok, and Neang, 2006, Raffles Bull. Zool., 54: 138.

Euphlyctis toumanoffi — Poynton and Broadley, 1985, Ann. Natal Mus., 27: 124, by implication.

Limnonectes (Bourretia) dabanus — Dubois, 1987 "1986", Alytes, 5: 62.

Limnonectes (Bourretia) toumanoffi — Dubois, 1987 "1986", Alytes, 5: 61–62.

Limnonectes (Elachyglossa) dabanus — Ohler and Dubois, 1999, Zool. Scripta, 23: 276.

Limnonectes (Elachyglossa) toumanoffi — Ohler and Dubois, 1999, Zool. Scripta, 23: 276.

English Names

Annam Wart Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 99).

Toumanoff's Wart Frog (Limnonectes toumanoffi [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 100).

Annam Fanged Frog (Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 23).

Distribution

Dak Lak, Dak Nong, Lam Dong, Dong Nai, Binh Dinh, Lam Dong, Ninh Thuan, and Khanh Hoa provinces, Vietnam, into adjacent Laos; tributaries of the Mekong and Krong Rivers in Cambodia and southern Vietnam, below 900 m elevation.

Geographic Occurrence

Natural Resident: Cambodia, Laos, Vietnam

Comment

Status elevated from subspecies under Limnonectes macrognathus by Dubois, 1987 "1986", Alytes, 5: 62; without discussion. Orlov, Murphy, Ananjeva, Ryabov, and Ho, 2002, Russ. J. Herpetol., 9: 87, commented on the range. Stuart, Sok, and Neang, 2006, Raffles Bull. Zool., 54: 137–138, provided a record and account for Cambodia. Account provided (as Rana toumanoffi) by Bourret, 1942, Batr. Indochine: 272–274. See Ohler and Dubois, 1999, Zool. Scripta, 23: 276, for reference to Vietnam record (as Limnonectes toumanoffi). Orlov, Murphy, Ananjeva, Ryabov, and Ho, 2002, Russ. J. Herpetol., 9: 87, commented on the range of nominal Limnonectes toumanoffi. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 23, provided specific localities for Vietnam for Limnonectes toumanoffi. See map, description of geographic range and habitat, and conservation status (as Limnonectes toumanoffi) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 484. Neang and Holden, 2008, Field Guide Amph. Cambodia: 89, provided a photograph, brief account of identification, ecology, and range in Cambodia. Rowley, Le, Hoang, and Altig, 2014, Zootaxa, 3881: 195–200, reported on advertisement call and larval morphology. Lambertz, Hartmann, Walsh, Geissler, and McLeod, 2014, Zool. J. Linn. Soc., 172: 117–132, compared the head ornamentation of Limnonectes dabanus, Limnonectes gyldenstolpei, Limnonectes macrognathus, and Limnonectes plicatellus and suggested that the name Elachoglossa be restricted to this clade. Teynié, Nguyen, Lorvelec, Piquet, Lottier, and David, 2014, Bull. Soc. Herpetol. France, 151: 21–52, provided the record for Laos. See account, photograph, and map for Vietnam in Vassilieva, Galoyan, Poyarkov, and Geissler, 2016, Photograph. Field Guide Amph. Rept. Lowland S. Vietnam: 90–92. Phimmachak, Richards, Sivongxay, Seateun, Chuaynkern, Makchai, Som, and Stuart, 2019, ZooKeys, 846: 133–156, provided morphological and molecular data in comparison with Limnonectes savan. Pham, Phan, Do, and Nguyen, 2019, Tap-Chi-Sinh-Hoc, 41 (2se1&2se2): 170–171, reported collection sites from Ninh Thuan and Khanh Hoa provinces, Vietnam, and described the specimens, provided photograph and ecological notes, and summarized the range in Vietnam by province.

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