Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Gastrotheca cornuta (Boulenger, 1898)

Class: Amphibia > Order: Anura > Family: Hemiphractidae > Genus: Gastrotheca > Species: Gastrotheca cornuta

Nototrema cornutum Boulenger, 1898, Proc. Zool. Soc. London, 1898: 124. Holotype: BMNH 1947.2.22.49 (formerly 1898.4.28.162) according to Condit, 1964, J. Ohio Herpetol. Soc., 4: 86. Type locality: "Cachabé" (= Cachabí), Provincia Esmeraldas, Ecuador. Noted elsewhere in the original publication as "Cachabé, a small village on the river of that name, on the N. W. Coast, in the Prov. Esmeraldas.... probably about 500 feet above the sea".

Hyla ceratophrys Stejneger, 1911, Proc. U.S. Natl. Mus., 41: 286. Holotype: USNM 47705, by original designation. Type locality: "upper Pequeni River, [Provincia Panamá,] Republic of Panama". Synonymy by Duellman, 1983, Copeia, 1983: 872.

Gastrotheca cornutum — Peters, 1955, Rev. Ecuat. Entomol. Parsitol., 2: 339.

Gastrotheca ceratophrys — Duellman, 1966, Univ. Kansas Publ. Mus. Nat. Hist., 17: 265.

Gastrotheca (Opisthodelphys) cornuta — Dubois, 1987 "1986", Alytes, 5: 31.

Gastrotheca (Amphignathodon) cornuta — Duellman, 2015, Marsupial Frogs: 163. Attributed to Dubois, 1987 "1986", in error.

Amphignathodon cornutus — Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 431.

Common Names

Horned Marsupial Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 53).

Distribution

Pacific lowlands of Ecuador, Colombia, and Panama; the Caribbean slopes of western Panama and adjacent Costa Rica, sea level to 1000 m elevation.

Geographic Occurrence

Natural Resident: Colombia, Costa Rica, Ecuador, Panama

Comment

In the Gastrotheca ovifera group of Duellman, Maxson, and Jesiolowski, 1988, Copeia, 1988: 527–543. See account (as Gastrotheca ceratophrys) by Duellman, 1970, Monogr. Mus. Nat. Hist. Univ. Kansas: 153–158. Duellman, 2001, Hylid Frogs Middle Am., Ed. 2: Tab. 67, treated Gastrotheca ceratophrys as a distinct species without discussion. See account by Savage, 2002, Amph. Rept. Costa Rica: 294. Köhler, 2011, Amph. Cent. Am.: 192–194, compared this species with other Central American hemiphractids and provided a map and photograph of this species. In the Gastrotheca longipes group of Castroviejo-Fisher, Padial, De la Riva, Pombal, Silva, Rojas-Runjaic, Medina-Méndez, and Frost, 2015, Zootaxa, 4004: 1–72. See account by Duellman, 2015, Marsupial Frogs: 163–169. Salazar-Zúñiga, Chaves-Acuña, Chaves, Acuña, Abarca-Odio, Lobón-Rovira, Gómez-Méndez, Gutiérrez-Vannucchi, and Bolaños, 2019, Amph. Rept. Conserv., 13 (2: e215): 304–322, first record in Panama in 21 years.

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to Ecuador see FaunaWebEcuador: Anfibios del Ecuador
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.