Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Gastrotheca riobambae (Fowler, 1913)

Class: Amphibia > Order: Anura > Family: Hemiphractidae > Genus: Gastrotheca > Species: Gastrotheca riobambae

Hyla riobambae Fowler, 1913, Proc. Acad. Nat. Sci. Philadelphia, 65: 157. Holotype: ANSP 18161, according to Malnate, 1971, Proc. Acad. Nat. Sci. Philadelphia, 123: corrigenda. Type locality: "Rio Bamba, [Provincia Chimborazo,] Ecuador".

Hyla quitoe Fowler, 1913, Proc. Acad. Nat. Sci. Philadelphia, 65: 159. Holotype: ANSP 18238 by original designation. Type locality: "Quito, [Provincia Pichincha,] Ecuador". Synonymy by Duellman and Fritts, 1972, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 9: 11.

Chorophilus olivaceus Andersson, 1945, Ark. Zool., 37A(2): 85. Holotype: NHRM 1965, according to Duellman, 1977, Das Tierreich, 95: 19. Type locality: "Rio Napo, 400 m"; supposed by Duellman, 1977, Das Tierreich, 95: 19, to be possibly "Rio Baños, Provincia Tunurahua, Ecuador". Synonymy by Duellman and Fritts, 1972, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 9: 11.

Gastrotheca marsupiata ecuatoriensis Vellard, 1957, Mem. Mus. Hist. Nat. Javier Prado, 5: 43. Nomen nudum. Synonymy by Duellman and Fritts, 1972, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 9: 11.

Gastrotheca riobambae — Duellman and Fritts, 1972, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 9: 11.

Gastrotheca cavia Duellman, 1974, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 22: 5. Holotype: KU 148532, by original designation. Type locality: "Isla Pequeña, Laguna Cuicocha, Provincia Imbabura, Ecuador, 2390 m". Synonymy by Duellman and Hillis, 1987, Herpetologica, 43: 163.

Gastrotheca (Duellmania) cavia — Dubois, 1987 "1986", Alytes, 5: 33.

Gastrotheca (Duellmania) riobambae — Dubois, 1987 "1986", Alytes, 5: 33; Duellman, 2015, Marsupial Frogs: 263.

Common Names

Riobamba Pouched Frog (Cochran, 1961, Living Amph. World: 141).

Riobamba Marsupial Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 53).

Distribution

Andes and inter-Andean valleys in northern and central Ecuador (Imbabura south to Chimborazo provinces), 2200–3220 m elevation.

Geographic Occurrence

Natural Resident: Ecuador

Endemic: Ecuador

Comment

In the Gastrotheca plumbea group of Duellman, Maxson, and Jesiolowski, 1988, Copeia, 1988: 527-543. See account by Duellman and Hillis, 1987, Herpetologica, 43: 159. Almendáriz C. and Orcés, 2004, Rev. Politécnica, Quito, 25: 117-118, provided distributional data for Ecuador. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 398. In the Gastrotheca marsupiata group of Castroviejo-Fisher, Padial, De la Riva, Pombal, Silva, Rojas-Runjaic, Medina-Méndez, and Frost, 2015, Zootaxa, 4004: 1–72. See detailed account by Duellman, 2015, Marsupial Frogs: 263–272. See comment by Reyes-Puig, Reyes-Puig, Franco-Mena, Jost, and Yánez-Muñoz, 2022, ZooKeys, 1081: 54, on the possibility of this species extending to Naturetrek–Vizcaya Reserve, Tunguruhua Province, Ecuador. Folly, Amaral, Silveira, Carvalho-e-Silva, and Dias, 2024, J. Morphol., 285(285:e21766): 1–12, reported on larval crania anatomy and discussed phylogenetic significance of these data.

External links:

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For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to Ecuador see FaunaWebEcuador: Anfibios del Ecuador
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.