Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Bokermannohyla Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005

Class: Amphibia > Order: Anura > Family: Hylidae > Genus: Bokermannohyla

30 species

Bokermannohyla Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 82. Type species: Hyla circumdata Cope, 1871 "1870", by original designation.

Common Names

Bokermann's Treefrogs (Faivovich, Lourenço, Pinheiro, Lyra, Baêta, Magalhães, Grant, Pezzuti, Leite, Araujo-Vieira, Brandão, Carvalho, Marinho, Orrico, Sabbag, Berneck, Pereyra, Giaretta, Toledo, Garcia, Pombal, Wheeler, Rodrigues, Napoli, and Haddad, 2025, Cladistics, 41: 323).

Distribution

Southern Brazil in the Distrito Federal and states of Paraná, Minas Gerais, Bahia, Goiás, Rio de Janeiro, São Paulo, and Santa Catarina.

Comment

This genus is composed of the former Hyla circumdata, Hyla martinsi, and Hyla pseudopseudis groups of previous authors. Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 83-84, recognized three species groups noted in the individual accounts, the Bokermannohyla circumdata group, Bokermannohyla claresignata group, Bokermannohyla martinsi group, and Bokermannohyla pseudopseudis group. In Cophomantini of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 75. See comments by Faivovich, Lugli, Lourenço, and Haddad, 2009, Herpetologica, 65: 303-310, on various diagnostic characters of species of Bokermannohyla. Leite and Eterovick, 2010, J. Herpetol., 44: 431-440, discussed aspects of comparative morphology of larvae. Catroli, Faivovich, Haddad, and Kasahara, 2011, J. Herpetol., 45: 120-128, reported on the remarkable homogeneity of karyotypes among three species groups of Bokermannohyla and Aplastodiscus and Hypsiboas. Pinheiro, Blotto, Ron, Stanley, Garcia, Haddad, Grant, and Faivovich, 2022, Zool. J. Linn. Soc., 195: 995–1021, provided an osteological synapomorphy for the Bokermannohyla martinsi group. Faivovich, Lourenço, Pinheiro, Lyra, Baêta, Magalhães, Grant, Pezzuti, Leite, Araujo-Vieira, Brandão, Carvalho, Marinho, Orrico, Sabbag, Berneck, Pereyra, Giaretta, Toledo, Garcia, Pombal, Wheeler, Rodrigues, Napoli, and Haddad, 2025, Cladistics, 41: 323–357, reported on phylogenetic relationships, biogeography, life history evolution, and redelimited species groups, noted in the individual species records. Pupin, Brusquetti, Leite, and Haddad, 2025, Zool. J. Linn. Soc., 205(1zlaf108): 1–14, reported on the molecular phylogenetics and biogeography of the Bokermannohyla pseudopseudis clade. Nakamura, Escalona Sulbarán, and Pinheiro, 2025 "2024", Biol. J. Linn. Soc., 144 (2: blae036): 1–18, reported on the drivers of call evolution.

Contained taxa (30 sp.):

Bokermannohyla ahenea (Napoli and Caramaschi, 2004)

Bokermannohyla alvarengai (Bokermann, 1956)

Bokermannohyla astartea (Bokermann, 1967)

Bokermannohyla capra Napoli and Pimenta, 2009

Bokermannohyla caramaschii (Napoli, 2005)

Bokermannohyla carvalhoi (Peixoto, 1981)

Bokermannohyla circumdata (Cope, 1871)

Bokermannohyla diamantina Napoli and Juncá, 2006

Bokermannohyla feioi (Napoli and Caramaschi, 2004)

Bokermannohyla flavopicta Leite, Pezzuti, and Garcia, 2012

Bokermannohyla gouveai (Peixoto and Cruz, 1992)

Bokermannohyla hylax (Heyer, 1985)

Bokermannohyla ibitiguara (Cardoso, 1983)

Bokermannohyla ibitipoca (Caramaschi and Feio, 1990)

Bokermannohyla itapoty Lugli and Haddad, 2006

Bokermannohyla izecksohni (Jim and Caramaschi, 1979)

Bokermannohyla juiju Faivovich, Lugli, Lourenço, and Haddad, 2009

Bokermannohyla langei (Bokermann, 1965)

Bokermannohyla lucianae (Napoli and Pimenta, 2003)

Bokermannohyla luctuosa (Pombal and Haddad, 1993)

Bokermannohyla martinsi (Bokermann, 1964)

Bokermannohyla nanuzae (Bokermann and Sazima, 1973)

Bokermannohyla oxente Lugli and Haddad, 2006

Bokermannohyla pseudopseudis (Miranda-Ribeiro, 1937)

Bokermannohyla ravida (Caramaschi, Napoli, and Bernardes, 2001)

Bokermannohyla sagarana Leite, Pezzuti, and Drummond, 2011

Bokermannohyla sapiranga Brandão, Magalhães, Garda, Campos, Sebben, and Maciel, 2012

Bokermannohyla saxicola (Bokermann, 1964)

Bokermannohyla sazimai (Cardoso and Andrade, 1982)

Bokermannohyla vulcaniae (Vasconcelos and Giaretta, 2005)

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