Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Boana benitezi (Rivero, 1961)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Boana > Species: Boana benitezi

Hyla benitezi Rivero, 1961, Bull. Mus. Comp. Zool., 126: 116. Holotype: MCZ 28564, by original designation. Type locality: "Caño Wanadi, Mt. Marahuaca, Territorio Amazonas", Venezuela.

Centrolenella pulidoi Rivero, 1968, Mem. Soc. Cienc. Nat. La Salle, 28: 312. Holotype: MCZ 72499, by original designation. Type locality: "Monte Duida, 2.000 pies, Territorio Amazonas, Venezuela". Synonymy by Barrio-Amorós, Rojas-Runjaic, and Señaris, 2019, Amph. Rept. Conserv., 13 (1: e180): 1.

Hypsiboas pulidoi — Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 87.

Hypsiboas benitezi — Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 87.

Boana benitezi — Dubois, 2017, Bionomina, 11: 28.

Boana pulidoi — Dubois, 2017, Bionomina, 11: 28.

Common Names

Benitez's Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 54).

Distribution

West of Sierra de Maigualida, in Cerro Duida, Cerro Marahuaka, Cerro Jaua, and Cerro Tapirapeco, in the western part of Guyana Highlands in Amazonas, Venezuela, and adjacent Brazil.

Geographic Occurrence

Natural Resident: Brazil, Venezuela

Comment

See Rivero, 1972 "1971", Caribb. J. Sci., 11: 181–193. See also Donnelly and Myers, 1991, Am. Mus. Novit., 3017: 11–14. Heyer, 1994, J. Herpetol., 28: 497-499, discussed Brazilian record. See comments by Gorzula and Señaris, 1999 "1998", Scient. Guaianae, 8: 27–28. Señaris and Ayarzagüena, 2002, J. Herpetol., 36: 635, reported the species from Cerro Jaua at 1600 m. In the Hypsiboas benitezi group of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 86–87. Barrio-Amorós and Brewer-Carias, 2008, Zootaxa, 1942: 28, restricted the range to exclude what is now known as Hypsiboas tepuianus. Centrolene pulidoi was first noted as a hylid by Priscilla Starrett pers. comm. to J. M. Savage In Duellman, 1985, in Frost (ed.), Amph. Species World: 84. Señaris and Rojas-Runjaic, 2020, in Rull and Carnaval (eds.), Neotrop. Divers. Patterns Process.: 571–632, commented on range and conservation status in the Venezuelan Guayana.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.