Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Dryophytes gratiosus (LeConte, 1856)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Dryophytes > Species: Dryophytes gratiosus

Hyla gratiosa LeConte, 1856, Proc. Acad. Nat. Sci. Philadelphia, 8: 146. Syntypes: Three specimens noted in description, of which ANSP 2089 was recorded as "holotype" by Malnate, 1971, Proc. Acad. Nat. Sci. Philadelphia, 123: 351, this choice substantiated by its formal designation as lectotype by Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 343. Cochran, 1961, Bull. U.S. Natl. Mus., 220: 54, noted USNM 5901 (6 specimens) as syntypes. The status of type(s) is confusing, see Caldwell, 1982, Cat. Am. Amph. Rept., 298: 1, and Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 343, for discussion. Type locality: "lower country of Georgia", considered by Stejneger and Barbour, 1933, Check List N. Am. Amph. Rept., Ed. 3: 35, to be "evidently one of the LeConte plantations in Floyd or Liberty County", Georgia, USA; restricted to "Liberty County, Georgia", USA, by Schmidt, 1953, Check List N. Am. Amph. Rept., Ed. 6: 71, although as not based on disclosed evidence this restriction is invalid according to Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 343.

Epedaphus gratiosus — Cope, 1885 "1884", Proc. Am. Philos. Soc., 22: 383.

Hyla (Epedaphus) gratiosa — Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 343.

Dryophytes gratiosus — Duellman, Marion, and Hedges, 2016, Zootaxa, 4104: 23.

English Names

Florida Hyla (Yarrow, 1882, Bull. U.S. Natl. Mus., 24: 24).

Florida Tree Frog (Dickerson, 1906, The Frog Book: 124; Wright, 1932, Life Hist. Frogs Okefinokee Swamp, 2: 14; Wright and Wright, 1933, Handb. Frogs Toads U.S. Canada: x).

Georgia Tree Frog (Brimley, 1907, J. Elisha Mitchell Sci. Soc., 23: 158).

Bell-frog (Carr, 1940, Univ. Florida Biol. Sci. Ser., 3: 60).

Giant Tree Frog (Wright and Wright, 1933, Handb. Frogs Toads U.S. Canada: x; Viosca, 1949, Pop. Sci. Bull., Louisiana Acad. Sci., 1: 10).

Barking Tree Frog (Schmidt, 1953, Check List N. Am. Amph. Rept., Ed. 6: 71).

Barking Treefrog (Conant, Cagle, Goin, Lowe, Neill, Netting, Schmidt, Shaw, Stebbins, and Bogert, 1956, Copeia, 1956: 176; Conant, 1975, Field Guide Rept. Amph. E. Cent. N. Am., Ed. 2: 321; Collins, Huheey, Knight, and Smith, 1978, Herpetol. Circ., 7: 11; Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 55; Collins, 1997, Herpetol. Circ., 25: 12; Crother, Boundy, Campbell, de Queiroz, Frost, Highton, Iverson, Meylan, Reeder, Seidel, Sites, Taggart, Tilley, and Wake, 2001 "2000", Herpetol. Circ., 29: 10; Frost, McDiarmid, and Mendelson, 2008, in Crother (ed.), Herpetol. Circ., 37: 6; Collins and Taggart, 2009, Standard Common Curr. Sci. Names N. Am. Amph. Turtles Rept. Crocodil., ed. 6: 7; Frost, McDiarmid, Mendelson, and Green, 2012, in Crother (ed.), Herpetol. Circ., 39: 15; Frost, Lemmon, McDiarmid, and Mendelson, 2017, in Crother (ed.), Herpetol. Circ., 43: 12).

Distribution

Southeastern USA (eastern Virginia to eastern Louisiana); isolated records from southwestern Kentucky, southern New Jersey, Delaware, Tennessee, and northern Alabama.

Geographic Occurrence

Natural Resident: United States of America, United States of America - Alabama, United States of America - Delaware, United States of America - Florida, United States of America - Georgia, United States of America - Kentucky, United States of America - Louisiana, United States of America - Maryland, United States of America - Mississippi, United States of America - New Jersey, United States of America - North Carolina, United States of America - South Carolina, United States of America - Tennessee, United States of America - Virginia

Endemic: United States of America

Comment

In the Hyla cinerea group of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 102. Reviewed by Caldwell, 1982, Cat. Am. Amph. Rept., 298: 1–2. Mitchell, 2005, in Lannoo (ed.), Amph. Declines: 455–456, and Dodd, 2013, Frogs U.S. and Canada, 1: 280–288, provided accounts that summarized the relevant literature. Elliot, Gerhardt, and Davidson, 2009, Frogs and Toads of N. Am.: 48–51, provided an account, photos, and advertisement call. Altig and McDiarmid, 2015, Handb. Larval Amph. US and Canada: 196–198, provided an account of larval morphology and biology. See account of biology and life history in southern Florida by Meshaka and Lane, 2015, Herpetol. Conserv. Biol., 10 (Monogr. 5): 54–56. Guyer and Bailey, 2023, Frogs and Toads of Alabama: 210–214, provided a detailed account for the species in Alabama, USA.

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.