Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Dryophytes Fitzinger, 1843

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Dryophytes

20 species

Dryophytes Fitzinger, 1843, Syst. Rept.: 31. Type species: Hyla versicolor LeConte, 1825, by original designation.

Epedaphus Cope, 1885 "1884", Proc. Am. Philos. Soc., 22: 383. Type species: Hyla gratiosa LeConte, 1856, by monotypy. Synonymy with Hyla by Kellogg, 1932, Bull. U.S. Natl. Mus., 160: 149. Treatment as a subgenus by Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 331–332. Synonymy with Dryophytes by implication of Duellman, Marion, and Hedges, 2016, Zootaxa, 4104: 23.

Dryophytes — Duellman, Marion, and Hedges, 2016, Zootaxa, 4104: 23.

Common Names

Ameroasian Treefrogs (Hedges, Powell, Henderson, Hanson, and Murphy, 2019, Caribb. Herpetol., 67: 13).

Holarctis Treefrogs (Guyer and Bailey, 2023, Frogs and Toads of Alabama: 183).

Distribution

North America east of the Sierra Nevada southward from extreme southern Canada to the Gulf of Mexico and on the Mexican Plateau southward to Oaxaca, plus the highlands of Chiapas, Mexico, and adjacent Guatemala; Far eastern Russia, Korean Peninsula, Japan, eastern China, and Ryukyu Island

Comment

See comments under Hyla for access to relevant literature. Removed from the synonymy of Hyla by Duellman, Marion, and Hedges, 2016, Zootaxa, 4104: 23 (who could find no morphological synapomorphies), where it had been placed by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 373. Previously, Dryophytes had been treated as a subgenus of Hyla by Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 331–332. On the basis of phylogenetic analysis of the mitogenome Zhang, Luu, Yu, Zhang, Al-attar, and Storey, 2019, Int. J. Biol. Macromolecules, 132: 461–469, suggested the relationship of Dyophytes versicolor + ((Dyophytes japonicus + Dyophytes ussuriensis) + (Dyophytes suweonensis + Dyophytes immaculatus)). Borzée, Messenger, Chae, Andersen, Groffen, Kim, An, Othman, Ri, Nam, Bae, Ren, Li, Chuang, Yi, Shin, Kwon, Jang, and Min, 2020, PLoS One, 15(6: e0234299): 1–34, revised the species of the Korean peninsula and delimited the Dryophtyes immaculatus group.

Contained taxa (20 sp.):

Dryophytes andersonii (Baird, 1854)

Dryophytes arboricola (Taylor, 1941)

Dryophytes arenicolor (Cope, 1866)

Dryophytes avivoca (Viosca, 1928)

Dryophytes bocourti (Mocquard, 1899)

Dryophytes chrysoscelis (Cope, 1880)

Dryophytes cinereus (Schneider, 1799)

Dryophytes euphorbiaceus (Günther, 1858)

Dryophytes eximius (Baird, 1854)

Dryophytes femoralis (Daudin, 1800)

Dryophytes flaviventris Borzée and Min, 2020

Dryophytes gratiosus (LeConte, 1856)

Dryophytes immaculatus (Boettger, 1888)

Dryophytes japonicus (Günther, 1859)

Dryophytes plicatus (Brocchi, 1877)

Dryophytes squirellus (Daudin, 1800)

Dryophytes suweonensis (Kuramoto, 1980)

Dryophytes versicolor (LeConte, 1825)

Dryophytes walkeri (Stuart, 1954)

Dryophytes wrightorum (Taylor, 1939)

External links:

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