Dryophytes Fitzinger, 1843

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Dryophytes
20 species

Dryophytes Fitzinger, 1843, Syst. Rept.: 31. Type species: Hyla versicolor LeConte, 1825, by original designation. 

Epedaphus Cope, 1885 "1884", Proc. Am. Philos. Soc., 22: 383. Type species: Hyla gratiosa LeConte, 1856, by monotypy. Synonymy with Hyla by Kellogg, 1932, Bull. U.S. Natl. Mus., 160: 149. Treatment as a subgenus  by Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 331–332.  Synonymy with Dryophytes by implication of Duellman, Marion, and Hedges, 2016, Zootaxa, 4104: 23. 

DryophytesDuellman, Marion, and Hedges, 2016, Zootaxa, 4104: 23. 

English Names

Ameroasian Treefrogs (Hedges, Powell, Henderson, Hanson, and Murphy, 2019, Caribb. Herpetol., 67: 13). 

Holarctis Treefrogs (Guyer and Bailey, 2023, Frogs and Toads of Alabama: 183).


North America east of the Sierra Nevada southward from extreme southern Canada to the Gulf of Mexico and on the Mexican Plateau southward to Oaxaca, plus the highlands of Chiapas, Mexico, and adjacent Guatemala; Far eastern Russia, Korean Peninsula, Japan, eastern China, and Ryukyu Island


See comments under Hyla for access to relevant literature. Removed from the synonymy of Hyla by Duellman, Marion, and Hedges, 2016, Zootaxa, 4104: 23 (who could find no morphological synapomorphies), where it had been placed by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 373. Previously, Dryophytes had been treated as a subgenus of Hyla by Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 331–332. On the basis of phylogenetic analysis of the mitogenome Zhang, Luu, Yu, Zhang, Al-attar, and Storey, 2019, Int. J. Biol. Macromolecules, 132: 461–469, suggested the relationship of Dyophytes versicolor + ((Dyophytes japonicusDyophytes ussuriensis) + (Dyophytes suweonensisDyophytes immaculatus)). Borzée, Messenger, Chae, Andersen, Groffen, Kim, An, Othman, Ri, Nam, Bae, Ren, Li, Chuang, Yi, Shin, Kwon, Jang, and Min, 2020, PLoS One, 15(6: e0234299): 1–34, revised the species of the Korean peninsula and delimited the Dryophtyes immaculatus group. 

Contained taxa (20 sp.):

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.