Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Isthmohyla lancasteri (Barbour, 1928)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Isthmohyla > Species: Isthmohyla lancasteri

Hyla lancasteri Barbour, 1928, Proc. New England Zool. Club, 10: 31. Holotype: MCZ 13062, by original designation. Type locality: "high forest at Peralta, [Cantón de Turrialba, Provincia Cartago,] Costa Rica, altitude 500 meters". Savage, 1974, Rev. Biol. Tropical, 22: 101, commented on the type locality.

Hyla moraviaensis Taylor, 1952, Univ. Kansas Sci. Bull., 35: 865. Holotype: KU 30284, by original designation. Type locality: "Moravia, [Cantón de Turrialba, Cartago Province,] Costa Rica (Caribbean drainage)". Savage, 1974, Rev. Biol. Tropical, 22: 98, commented on the type locality. Synonymy by Duellman, 1966, Univ. Kansas Publ. Mus. Nat. Hist., 17: 271.

Isthmohyla lancasteri — Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 103.

Common Names

Lancaster's Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 56).

Distribution

Humid premontane slopes of the Cordillera de Talamanca of Costa Rica and western Panama, 40–1450 m elevation.

Geographic Occurrence

Natural Resident: Costa Rica, Panama

Comment

See Myers and Duellman, 1982, Am. Mus. Novit., 2752: 15, and Duellman, 2001, Hylid Frogs Middle Am., Ed. 2: 878–879, who provided an account and transferred the former Hyla lancasteri group into the Hyla pictipes group. See comment under Hyla calypsa with which this was formerly confused. Lips and Savage, 1996, Proc. Biol. Soc. Washington, 109: 17–26, included this species (as Hyla lancasteri) in a key to the tadpoles found in Costa Rica. See account by Savage, 2002, Amph. Rept. Costa Rica: 311–312. In the Isthmohyla pictipes group of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 103. Köhler, 2011, Amph. Cent. Am.: 236–244, provided a brief summary of natural history for the species of Isthmohyla in Central America and provided a range map and photograph for this species. Faivovich, Pereyra, Luna, Hertz, Blotto, Vásquez-Almazán, McCranie, Sánchez, Baêta, Araujo-Vieira, Köhler, Kubicki, Campbell, Frost, Wheeler, and Haddad, 2018, S. Am. J. Herpetol., 13: 16, discussed the need for further taxonomic elucidation of this nominal species, which might contain at least one other lineage.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.