Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Isthmohyla Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Isthmohyla

13 species

Isthmohyla Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 102. Type species: Hyla pseudopuma Günther, 1901, by original designation.

Common Names

None noted.

Distribution

Honduras, Costa Rica, and Panama.

Comment

As originally formed this taxon contains all members of the former Hyla pseudopuma and Hyla pictipes group as defined by Duellman, 2001, Hylid Frogs Middle Am., Ed. 2, with the exception of Hyla thorectes, which was transferred into Plectrohyla. Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 98, placed Isthmohyla in their Hylini and recognized two species groups: Isthmohyla pseudopuma group and Isthmohyla pictipes group (formed by union of the former Hyla lancasteri, Hyla pictipes, Hyla rivularis, and Hyla zeteki groups of Duellman, 1970, Monogr. Mus. Nat. Hist. Univ. Kansas). See comment under "Hyla" melacaena which likely is a member of this taxon. Faivovich, Pereyra, Luna, Hertz, Blotto, Vásquez-Almazán, McCranie, Sánchez, Baêta, Araujo-Vieira, Köhler, Kubicki, Campbell, Frost, Wheeler, and Haddad, 2018, S. Am. J. Herpetol., 13: 15–18, discussed the genus and its intrarelationships, including discussions of the species groups and phylogenetic ambiguities.

Contained taxa (13 sp.):

Isthmohyla angustilineata (Taylor, 1952)

Isthmohyla calypsa (Lips, 1996)

Isthmohyla debilis (Taylor, 1952)

Isthmohyla graceae (Myers and Duellman, 1982)

Isthmohyla infucata (Duellman, 1968)

Isthmohyla lancasteri (Barbour, 1928)

Isthmohyla picadoi (Dunn, 1937)

Isthmohyla pictipes (Cope, 1875)

Isthmohyla pseudopuma (Günther, 1901)

Isthmohyla rivularis (Taylor, 1952)

Isthmohyla tica (Starrett, 1966)

Isthmohyla xanthosticta (Duellman, 1968)

Isthmohyla zeteki (Gaige, 1929)

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