Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Sarcohyla labeculata (Shannon, 1951)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Sarcohyla > Species: Sarcohyla labeculata

Hyla bistincta labeculata Shannon, 1951, Proc. U.S. Natl. Mus., 101: 470. Holotype: USNM 123689, by original designation. Type locality: "San Lucas Camotlán, Oaxaca", Mexico.

Hyla calthula Ustach, Mendelson, McDiarmid, and Campbell, 2000, Herpetologica, 56: 239. Holotype: UTA A-5885, by original designation. Type locality: "Mexico: Oaxaca: Sierra Mixes, Totontepec". Synonymy by Campbell, Brodie, Caviedes-Solis, Nieto-Montes de Oca, Luja, Flores-Villela, García-Vázquez, Sarker, and Wostl, 2018, Zootaxa, 4422: 373.

Plectrohyla calthula — Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 105.

Hyla ephemera Meik, Canseco-Márquez, Smith, and Campbell, 2005, Zootaxa, 1046: 20. Holotype: MZFC 17049, by original designation. Type locality: "between Santa Maria Guienagati and Lachidola, 1100 m, (16.759° N 95.461° W), north slope of Cerro Las Flores, Oaxaca, Mexico". Synonymy by Campbell, Brodie, Caviedes-Solis, Nieto-Montes de Oca, Luja, Flores-Villela, García-Vázquez, Sarker, and Wostl, 2018, Zootaxa, 4422: 373.

Sarcohyla calthula — Duellman, Marion, and Hedges, 2016, Zootaxa, 4104: 18.

Sarcohyla ephemera — Duellman, Marion, and Hedges, 2016, Zootaxa, 4104: 18.

Sarcohyla labeculata — Campbell, Brodie, Caviedes-Solis, Nieto-Montes de Oca, Luja, Flores-Villela, García-Vázquez, Sarker, and Wostl, 2018, Zootaxa, 4422: 373.

English Names

Mixe Streamside Treefrog (Campbell, Brodie, Caviedes-Solis, Nieto-Montes de Oca, Luja, Flores-Villela, García-Vázquez, Sarker, and Wostl, 2018, Zootaxa, 4422: 373).

Yellow-robed Treefrog (Sarcohyla calthulua, no longer recognized: Liner and Casas-Andreu, 2008, Herpetol. Circ., 38: 19).

Cerro Las Flores Spikethumb Frog (Sarcohyla ephemera, no longer recognized: Liner and Casas-Andreu, 2008, Herpetol. Circ., 38: 20).

Distribution

The Sierra Mixe in the north-central Oaxacan highlands where it is known from along small streams and seeps in cloud forest and secondary growth near the village of Totontepec at 1780–1896 m, and also west of the village of Zacatepec (17.190° N, 95.987° W) at an elevation of 1360 m; an isolated population occurs on Cerro Las Flores, an uplift somewhat isolated from the rest of the Sierra Mixe and about 90 km southeast of Totontepec, where it has been taken in cloud forest at 1100–1220 m.

Comment

Resurrected from the synonymy of Sarcohyla bistincta by where it had been placed by Duellman, 1964, Univ. Kansas Publ. Mus. Nat. Hist., 15: 475. See accounts as Hyla calthula by Duellman, 2001, Hylid Frogs Middle Am., Ed. 2: 964-966. See illustration, map, description of geographic range and habitat, and conservation status as Plectrohyla calthula and Plectorhyla ephemera in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 267, 270.

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