Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Tepuihyla tuberculosa (Boulenger, 1882)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Tepuihyla > Species: Tepuihyla tuberculosa

Hyla tuberculosa Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 355. Holotype: BMNH 1947.2.13.34 (formerly 1880.12.5.172) according to Condit, 1964, J. Ohio Herpetol. Soc., 4: 95; presumably this is the animal figured in plate 24, fig. 1 of the original publication. Type locality: "Canelos", Provincia Pastaza, Ecuador.

Ecnomiohyla tuberculosa — Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 100. Provisional arrangement.

Hypsiboas tuberculosa — Duellman, Marion, and Hedges, 2016, Zootaxa, 4104: 21. Provisional placement.

Tepuihyla tuberculosa — Ron, Venegas, Ortega-Andrade, Gagliardi-Urrutia, and Salerno, 2016, ZooKeys, 630: 127.

Boana tuberculosa — Dubois, 2017, Bionomina, 11: 28.

Common Names

Canelos Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 58).

Distribution

Upper Amazon Basin in Ecuador and northeastern Peru, 132 to 1076 m elevation. Record for Amazonas, Brazil, presumably applies to Tepuihyla shushupe; record from southeastern Colombia presumably applies to Tepuihyla shushupe as well and requires confirmation.

Geographic Occurrence

Natural Resident: Ecuador, Peru

Likely/Controversially Present: Brazil, Colombia

Comment

See Duellman, 1974, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 27: 1–27 Rodríguez and Duellman, 1994, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 22: 38–39, provided a brief account for the Iquitos region of northeastern Peru as Hyla tuberculosa. Savage and Kubicki, 2010, Zootaxa, 2719: 22, regarded the assignment of this species to Ecnomiohyla as problematic as it lacks the morphological synapmorphies they associate with application of the generic name. Forti, Foratto, Márquez, Pereira, and Toledo, 2018, PeerJ, 6(e4813): 1–19, characterized the advertisement call. See discussion by Duellman, Marion, and Hedges, 2016, Zootaxa, 4104: 21, on why they provisionally placed this species in Hypsiboas. Ron, Venegas, Ortega-Andrade, Gagliardi-Urrutia, and Salerno, 2016, ZooKeys, 630: 115–154, provided an extensive discussion of the systematics and identification of the species and demonstrated that former Ecnomiohyla tuberculosa is a member of Tepuihyla. Gagliardi-Urrutia, García Dávila, Jaramillo-Martinez, Rojas-Padilla, Rios-Alva, Aguilar-Manihuari, Pérez-Peña, Castroviejo-Fisher, Simões, Estivals, Guillen Huaman, Castro Ruiz, Angulo Chávez, Mariac, Duponchelle, and Renno, 2022, Anf. Loreto: 130–131, provided a brief account, photograph, dot map, and genetic barcode for Loreto, Peru.

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to Ecuador see FaunaWebEcuador: Anfibios del Ecuador
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.