Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Tepuihyla Ayarzagüena, Señaris, and Gorzula, 1993

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Tepuihyla

9 species

Tepuihyla Ayarzagüena, Señaris Vasquez, and Gorzula, 1993 "1992", Mem. Soc. Cienc. Nat. La Salle, 52: 213. Type species: Hyla rodriguezi Rivero, 1968, by original designation.

Common Names

Tepui Treefrogs (Kok and Kalamandeen, 2008, Intr. Taxon. Amph. Kaieteur Natl. Park: 200-202; Coloma and Duellman, 2025, Amph. Ecuador. Vol. 2: 519).

Amazon Treefrogs (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 65).

Distribution

Mountains of eastern and southeastern Venezuela and adjacent Guyana and Brazil to the upper Amazon Basin of Ecuador and Peru.

Comment

This genus conforms to the Osteocephalus rodriguezi group as discussed by Ayarzagüena, Señaris Vasquez, and Gorzula, 1993 "1992", Mem. Soc. Cienc. Nat. La Salle, 52: 113-142, who discussed the group (as the Osteocephalus rodriguezi group) in the Venezuelan Guyana region, named five new species, and provided a key. In Lophyohylini of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 107-108. Myers and Stothers, 2006, Arch. Nat. Hist., London, 33: 241-266, discussed why the gender of this generic name must be considered feminine in spite of the stated intention of the original authors. Kok, MacCulloch, Means, Roelants, Van Bocxlaer, and Bossuyt, 2012, Curr. Biol., 22: R589-R590, provided a molecular tree of species as did Salerno, Ron, Señaris Vasquez, Rojas-Runjaic, Noonan, and Cannatella, 2012, Evolution, 66: 3000–3013. Salerno, Señaris Vasquez, Rojas-Runjaic, and Cannatella, 2015, Mol. Phylogenet. Evol., 82: 314–323, reported on the molecular phylogenetics of the Tepui species of Venezuela. Blotto, Lyra, Cardoso, Rodrigues, Dias, Marciano, Vechio, Orrico, Brandão, Assis, Lantyer-Silva, Rutherford, Gagliardi-Urrutia, Solé, Baldo, Nunes, Cajade, Torres, Grant, Jungfer, Silva, Haddad, and Faivovich, 2021, Cladistics, 37: 36–72, reported on molecular phylogenetics within the species and among the genera of lophiohyline hylids. Ortiz, Hoskin, Werneck, Réjaud, Manzi, Ron, and Fouquet, 2023, Organisms Divers. Evol., 23: 395–414, reported on the historical biogeography of Amazonia based on molecular evidence for Osteocephalus, Tepuihyla, and Dryaderces.

Contained taxa (9 sp.):

Tepuihyla aecii (Ayarzagüena, Señaris, and Gorzula, 1993)

Tepuihyla edelcae (Ayarzagüena, Señaris, and Gorzula, 1993)

Tepuihyla exophthalma (Smith and Noonan, 2001)

Tepuihyla luteolabris (Ayarzagüena, Señaris, and Gorzula, 1993)

Tepuihyla obscura Kok, Ratz, Tegelaar, Aubret, and Means, 2015

Tepuihyla rodriguezi (Rivero, 1968)

Tepuihyla shushupe Ron, Venegas, Ortega-Andrade, Gagliardi-Urrutia, and Salerno, 2016

Tepuihyla tuberculosa (Boulenger, 1882)

Tepuihyla warreni (Duellman and Hoogmoed, 1992)

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