Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Tepuihyla warreni (Duellman and Hoogmoed, 1992)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Tepuihyla > Species: Tepuihyla warreni

Hyla warreni Duellman and Hoogmoed, 1992, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 147: 10. Holotype: BMNH 1979.561, by original designation. Type locality: "north slope of Mt. Roraima (05° 38′ N, 60° 44′ W, elev. 1,480 m), Rupununi District, Guyana". Warren, 1973, Roraima: 47, mapped the locality (discussed on page 80) as ca. 05°17′ N.

Tepuihyla warreni — Aubrecht, Barrio-Amorós, Breure, Brewer-Carias, Derka, Fuentes-Ramos, Gregor, Kodada, Kováčik, Lánczos, Lee, Liščák, Schlögl, Šmída, and Vlček, 2012, Venezuelan Tepuis: XXX. Unexplained combination.

Tepuihyla warreni — Jungfer, Faivovich, Padial, Castroviejo-Fisher, Lyra, Berneck, Iglesias, Kok, MacCulloch, Rodrigues, Verdade, Torres-Gastello, Chaparro, Valdujo, Reichle, Moravec, Gvoždík, Gagliardi-Urrutia, Ernst, De la Riva, Means, Lima, Señaris, Wheeler, and Haddad, 2013, Zool. Scripta, 42 : 157.

English Names

Warren's Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 58).

Distribution

Mount Roraima and Mount Ayanganna in southwestern Guyana.

Geographic Occurrence

Natural Resident: Guyana

Endemic: Guyana

Comment

Not assignable to species group, according to the original publication. Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 111, could not assign this species to any of the currently recognized taxa, but pending examination of the type left this species incertae sedis. MacCulloch and Lathrop, 2005, Phyllomedusa, 4: 24–25, reported the species on Mt. Ayanganna. Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294, could not allocate several members of former "Hyla" to any of the new genera but believed that they would be so allocated with additional work. As an interim measure they merely allocated these species to the non-taxon "Hyla", clearly not part of the North American-Eurasian clade which bears the name Hyla. MacCulloch and Lathrop, 2009, R. Ontario Mus. Contrib. Sci., 4: 13, commented on specimens from Mount Ayanganna, Guyana. Kok, MacCulloch, Means, Roelants, Van Bocxlaer, and Bossuyt, 2012, Curr. Biol., 22: R589-R590, and Kok, MacCulloch, Means, Roelants, Van Bocxlaer, and Bossuyt, 2012, Current. Biol., Suppl. Inform., 22: 1-25, provided a molecular tree of tepui populations that suggested that this species may be associated with Tepuihyla, a not-so-surprising result. See Cole, Townsend, Reynolds, MacCulloch, and Lathrop, 2013, Proc. Biol. Soc. Washington, 125: 402, for brief account and records for Guyana. Jungfer, Faivovich, Padial, Castroviejo-Fisher, Lyra, Berneck, Iglesias, Kok, MacCulloch, Rodrigues, Verdade, Torres-Gastello, Chaparro, Valdujo, Reichle, Moravec, Gvoždík, Gagliardi-Urrutia, Ernst, De la Riva, Means, Lima, Señaris, Wheeler, and Haddad, 2013, Zool. Scripta, 42 : 357, noted that this species is the sister taxon of Tepuihyla and therefore transferred the species formally into Tepuihyla on the basis of molecular analysis.

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For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.