Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Phyllomedusa tetraploidea Pombal and Haddad, 1992

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Phyllomedusinae > Genus: Phyllomedusa > Species: Phyllomedusa tetraploidea

Phyllomedusa tetraploidea Pombal and Haddad, 1992, Rev. Brasil. Biol., 52: 219. Holotype: ZUEC 7589, by original designation. Type locality: "Holambra II, município de Paranapanema (approx. 23° 23′ S, 48° 44′ W), Estado de São Paulo", Brazil.

Common Names

Hidden Walking Leaf Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 62).

Distribution

Interior regions of the states of São Paulo and Paraná, Brazil; Misiones, Argentina; extreme southeastern Paraguay (Itapúa).

Geographic Occurrence

Natural Resident: Argentina, Brazil, Paraguay

Comment

In the Phyllomedusa burmeisteri group, and derived by tetraploidy from Phyllomedusa distincta, according to the original publication. See Langone and Carrizo, 1996, Cuad. Herpetol., 10: 58-61, for Argentinian record. Lavilla and Cei, 2001, Monogr. Mus. Reg. Sci. Nat. Torino, 28: 62, noted that Argentinian records of Phyllomedusa iheringii apply to this species. In the Phyllomedusa burmeisteri group of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 117, and Faivovich, Haddad, Baêta, Jungfer, Álvares, Brandão, Sheil, Barrientos, Barrio-Amorós, Cruz, and Wheeler, 2010, Cladistics, 26: 259. Brusquetti and Lavilla, 2006, Cuad. Herpetol., 20: 13, provided the first record for Paraguay. Brunes, Sequeira, Haddad, and Alexandrino, 2010, Mol. Phylogenet. Evol., 57: 1120-1133, suggested on the basis of molecular analysis that this species either is derived as a tetraploid from Phyllomedusa distincta or from the common ancestor. Weiler, Núñez, Airaldi, Lavilla, Peris, and Baldo, 2013, Anf. Paraguay: 72, provided a brief account, image, and dot map for Paraguay. Röhr, Camurugi, Paterno, Gehara, Juncá, Álvares, Brandão, and Garda, 2020, Canad. J. Zool., 98: 495–504, reported on the evolution and causes of variability of advertisement call.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.