Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hyperolius balfouri (Werner, 1908)

Class: Amphibia > Order: Anura > Family: Hyperoliidae > Subfamily: Hyperoliinae > Genus: Hyperolius > Species: Hyperolius balfouri

Rappia balfouri Werner, 1908 "1907", Sitzungsber. Akad. Wiss. Wien, Phys. Math. Naturwiss. Kl., 116: 1904. Syntypes: including NHMW 22894, according to Häupl and Tiedemann, 1978, Kat. Wiss. Samml. Naturhist. Mus. Wien, 2: 28, and Häupl, Tiedemann, and Grillitsch, 1994, Kat. Wiss. Samml. Naturhist. Mus. Wien, 9: 33. Type locality: "Gondokoro", Uganda.

Hyperolius balfouri — Noble, 1924, Bull. Am. Mus. Nat. Hist., 49: 251, 324; Laurent, 1943, Ann. Mus. R. Congo Belge, Tervuren, Zool., 4: 75.

Hyperolius zavattarii Scortecci, 1943, in Zavattari (ed.), Miss. Biol. Sagan–Omo, 7(Zool. 1): 319. Holotype: MSNM Am 284 according to Blackburn and Scali, 2014, Herpetol. Monogr., 28: 27, who discussed the material.. Type locality: "Pozzi di Gondaraba", Ethiopia. Synonymy by Largen, 1998, Tropical Zool., 11: 75.

Hyperolius concolor balfouri — Laurent, 1950, Explor. Parc Natl. Albert, Miss. G.F. de Witte (1933–1935), 64: 57.

Hyperolius concolor viridistriatus Monard, 1951, Mem. Inst. Franç. Afr. Noire, 1: 176. Syntypes: LCFM (10 specimens). Type locality: "Tibati" and "Sakdjé", Cameroon. Synonymy with Hyperolius concolor by Schiøtz, 1967, Spolia Zool. Mus. Haun., 25: 179, with Rappia balfouri by Schiøtz, 1975, Treefrogs E. Afr.: 115.

Hyperolius viridistriatus — Perret, 1966, Zool. Jahrb., Jena, Abt. Syst., 93: 414-415.

Hyperolius balfouri balfouri — Schiøtz, 1975, Treefrogs E. Afr.: 113.

Hyperolius balfouri viridistriatus — Schiøtz, 1975, Treefrogs E. Afr.: 113.

English Names

Balfour's Reed Frog (Hyperolius balfouri: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 66; Channing and Howell, 2006, Amph. E. Afr.: 154).

Ethiopia Reed Frog (Hyperolius zavattarii [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 68).

Distribution

Cameroon to Gabon and east through Central African Republic, northern Rep. Congo, northern Dem. Rep. Congo and South Sudan to Uganda and southwestern Ethiopia and western Kenya, possibly into southern Chad and southeastern Nigeria.

Comment

In the Hyperolius concolor superspecies and including Hyperolius viridistriatus, according to Schiøtz, 1975, Treefrogs E. Afr.: 112. See account (as Hyperolius concolor balfouri) by Inger, 1968, Explor. Parc Natl. Garamba, Miss. H. de Saeger, 52: 144. See account of Ethiopian population by Largen, 1998, Tropical Zool., 11: 61–82. Schiøtz, 1999, Treefrogs Afr.: 128–129, who provided a brief account, suggested that the subspecies might be unwarranted. Largen, 2001, Tropical Zool., 14: 358, commented on distribution. Channing and Howell, 2006, Amph. E. Afr.: 154, provided an account for East Africa. Schiøtz, 2006, Alytes, 24: 40-60, reported the species from Bokuma in central Dem. Rep. Congo. Jackson and Blackburn, 2007, Salamandra, 43: 149–164, provided a record for Nouabale-Ndoki National Park, Rep. Congo. Jacobsen, 2009, Afr. Herp News, 47: 2-20, reported this species from east-north-central Central African Republic. See account, photograph, and map for Ethiopia by Largen and Spawls, 2010, Amph. Rept. Ethiopia Eritrea: 112–113. See account for Cameroon by Amiet, 2012, Rainettes Cameroun: 271–277 Jongsma, Tobi, Dixon-MacCallum, Bamba-Kaya, Yoga, Mbega, Mve Beh, Emrich, and Blackburn, 2017, Amph. Rept. Conserv., 11 (1: e144): 1, provided a record from Haut-Ogooué province in southeastern Gabon. Dewynter and Frétey, 2019, Cah. Fondation Biotope, 27: 23, summarized the literature for Gabon. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 172–173, provided a brief account, photograph, and range map. See Portik, Bell, Blackburn, Bauer, Barratt, Branch, Burger, Channing, Colston, Conradie, Dehling, Drewes, Ernst, Greenbaum, Gvoždík, Harvey, Hillers, Hirschfeld, Jongsma, Kielgast, Kouete, Lawson, Leaché, Loader, Lötters, van der Meijden, Menegon, Müller, Nagy, Ofori-Boateng, Ohler, Papenfuss, Rößler, Sinsch, Rödel, Veith, Vindum, Zassi-Boulou, and McGuire, 2019, Syst. Biol., 68: 859–875 (supplemental fig. 1) for phylogenetic placement.

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