Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hyperolius benguellensis (Barboza du Bocage, 1893)

Class: Amphibia > Order: Anura > Family: Hyperoliidae > Subfamily: Hyperoliinae > Genus: Hyperolius > Species: Hyperolius benguellensis

Rappia benguellensis Barboza du Bocage, 1893, J. Sci. Math. Phys. Nat., Lisboa, Ser. 2, 3 (10): 119. Syntypes: MBL 17.220–223, according to Perret, 1976, Arq. Mus. Bocage, Ser. 2, 6: 26; destroyed in 1978 fire, according to Laurent, 1985, in Frost (ed.), Amph. Species World: 207. Type locality: "Cahata" = Caota, Benguella, Angola.

Rappia dombeensis Tornier, 1896, in Möbius (ed.), Deutsch Ost-Afr., 3: 150. Holotype: ZMB 6465, according to Tillack, de Ruiter, and Rödel, 2021, Zoosyst. Evol., Berlin, 97: 440. Type locality:”Dombe” = Dombe Grande, Benguela Province, Angola. Provisional placement in this synonymy; see comment.

Rappia oxyrhynchus Boulenger, 1901, Ann. Mus. R. Congo Belge, Tervuren, Zool., 2: 5. Syntypes: MRAC and BMNH (1947.2.9.49 [formerly 1901.3.12.26] and 1947.2.9.50–51 [formerly 1901.3.26.7–8] according to museum records). Laurent, 1943, Ann. Mus. R. Congo Belge, Tervuren, Zool., 4: 73, regarded MRAC 151, apparently in error, as the holotype. Type locality: "Pweto et de Lofoï", Dem. Rep. Congo. Synonymy with Hyperolius nasutus by Laurent, 1957, Explor. Parc Natl. Upemba, Miss. G.F. de Witte, 42: 3–47. See Channing, Moyer, and Burger, 2002, Afr. Zool., 37: 97. Placed in this synonymy by Channing, Hillers, Lötters, Rödel, Schick, Conradie, Rödder, Mercurio, Wagner, Dehling, Du Preez, Kielgast, and Burger, 2013, Zootaxa, 3620: 317.

Hyperolius oxyrhynchus—Noble, 1924, Bull. Am. Mus. Nat. Hist., 49: 252, 327; Laurent, 1943, Ann. Mus. R. Congo Belge, Tervuren, Zool., 4: 73.

Hyperolius benguellensis — Noble, 1924, Bull. Am. Mus. Nat. Hist., 49: 251, 324; Ahl, 1931, Das Tierreich, 55: 305.

Common Names

Benguella Long Reed Frog (Channing, Hillers, Lötters, Rödel, Schick, Conradie, Rödder, Mercurio, Wagner, Dehling, Du Preez, Kielgast, and Burger, 2013, Zootaxa, 3620: 317).

Bocage's Sharp-nosed Reed Frog (Lambiris, 1990 "1989", Monogr. Mus. Reg. Sci. Nat. Torino, 10: 172; Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 244).

Benguella Reed Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 66).

Benguela Reed Frog (Marques, Ceríaco, Blackburn, and Bauer, 2018, Proc. California Acad. Sci., Ser. 4, 65 (Suppl. II): 94).

Distribution

Western to southeastern Angola, northern Namibia, and northern Botswana; possibly to be found more widely but confused with Hyperolius nasutus.

Geographic Occurrence

Natural Resident: Angola, Botswana, Namibia

Comment

In the Hyperolius nasutus group and removed from the synonymy of Hyperolius nasutus by Amiet, 2005, Rev. Suisse Zool., 112: 271–310 (although on page 292, he allowed that it may be a synonym of Hyperolius nasutus), where it had been placed by Loveridge, 1953, Bull. Mus. Comp. Zool., 110: 362–363; and Schiøtz, 1999, Treefrogs Afr.: 97. See comment under Hyperolius viridis. Channing, Hillers, Lötters, Rödel, Schick, Conradie, Rödder, Mercurio, Wagner, Dehling, Du Preez, Kielgast, and Burger, 2013, Zootaxa, 3620: 317–319, provided an account, including a description of the advertisement call, and redelimited the species. Marques, Ceríaco, Blackburn, and Bauer, 2018, Proc. California Acad. Sci., Ser. 4, 65 (Suppl. II): 94–96, provided a map for Angola and a brief account. Not addressed by Portik, Bell, Blackburn, Bauer, Barratt, Branch, Burger, Channing, Colston, Conradie, Dehling, Drewes, Ernst, Greenbaum, Gvoždík, Harvey, Hillers, Hirschfeld, Jongsma, Kielgast, Kouete, Lawson, Leaché, Loader, Lötters, van der Meijden, Menegon, Müller, Nagy, Ofori-Boateng, Ohler, Papenfuss, Rößler, Sinsch, Rödel, Veith, Vindum, Zassi-Boulou, and McGuire, 2019, Syst. Biol., 68: 859–875. Baptista, António, and Branch, 2019, Amph. Rept. Conserv., 13 (e203): 96–130, reported on specimens from Bicuar National Park, southwestern Angola, and comment on the state of systematics. Tillack, de Ruiter, and Rödel, 2021, Zoosyst. Evol., Berlin, 97: 440, discussed the collector and collection as well as relevant correspondence surrounding the naming of Rappia dombeensis. They placed it in the Hyperolius nasutus complex, but without genetic confirmation did not think further assignment was possible; it is here included in this synonymy solely on the basis of proximity to the type locality of Hyperolius benguellensis, although this is just an attempt to bring more attention to the taxon rather than abandon it to incertae sedis (DRF). Conradie, Keates, Verburgt, Baptista, and Harvey, 2023, Amph. Rept. Conserv., 17: 31, provided an account and dot map of the species in Angola.

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For additional information see AmphibiaWeb report
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For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.