Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hyperolius guttatus Peters, 1875

Class: Amphibia > Order: Anura > Family: Hyperoliidae > Subfamily: Hyperoliinae > Genus: Hyperolius > Species: Hyperolius guttatus

Hyperolius guttatus Peters, 1875, Monatsber. Preuss. Akad. Wiss. Berlin, 1875: 207. Syntypes: ZMB 8378, 4489; ZMB 8378 considered lectotype by Laurent, 1961, Rev. Zool. Bot. Afr., 64: 73, and Perret, 1975, Rev. Suisse Zool., 82: 190. Lectotype redescribed by Frétey, Dubois, and Ohler, 2014, Alytes, 30: 20–21. Type localities: "Cameruns" and "Boutry", Cameroon; restricted by lectotype designation to "Cameruns". Corrected to the vicinity of Douala, Cameroon by Frétey, Dubois, and Ohler, 2014, Alytes, 30: 17, who discussed the types and type locality and redescribed the lectotype. Placement as a subspecies of Hyperolius ocellatus by Laurent, 1956, Rev. Zool. Bot. Afr., 53: 235. Synonymy with Hyperolius steindachneri by Perret, 1975, Rev. Suisse Zool., 82: 190. Considered possibly conspecific with Hyperolius ocellatus by Lötters, Gossmann, Obame, and Böhme, 2001, Herpetofauna, Weinstadt, 23: 29.

Rappia sordida Fischer, 1888, Jahrb. Hamburg. Wiss. Anst., 5: 10. Syntypes: in "Lübecker Museum", by original designation; including BMNH 1889.12.16.160–161 (now 1947.2.9.25–26 by museum records), according to Parker, 1936, Proc. Zool. Soc. London, 1936: 151 (who provided a redescription). Type locality: "Kamerun". Synonymy by Loveridge, 1938, Proc. New England Zool. Club, 17: 49–74; Perret, 1966, Zool. Jahrb., Jena, Abt. Syst., 93: 411. Nečas, Kielgast, Chinemerem, Rödel, Dolinay, and Gvoždík, 2024, Zool. J. Linn. Soc., 202(4): 13 (Zoobank Publication registration: 004579A5-F55C-4429-97C6-A52A23F4501D, 15 May 2024) placed it as a synonym of Hyperolius guttatus, but suggested that further study might render this a synonym of Hyperolius ibadanensis.

Rappia guttata — Tornier, 1896, in Möbius (ed.), Deutsch Ost-Afr., 3: 142.

Hyperolius sordidus — Noble, 1924, Bull. Am. Mus. Nat. Hist., 49: 252, 329; Ahl, 1931, Das Tierreich, 55: 335.

Hyperolius pulcher Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 48. Holotype: ZMB unnumbered, according to the original publication; ZMB 36088, according to Laurent, 1961, Rev. Zool. Bot. Afr., 64: 78. Holotype redescribed by Frétey, Dubois, and Ohler, 2014, Alytes, 30: 25–26. Type locality: "Japoma [near Douala], Kamerun". Synonymy with Hyperolius concolor by Laurent, 1961, Rev. Zool. Bot. Afr., 64: 78. The description in Ahl, 1931, Das Tierreich, 55: 308, appeared subsequently according to unpublished notes by Tillack, de Ruiter, and Rödel, 2021, Zoosyst. Evol., Berlin, 97: 432. Synonymy by implication of Frétey, Dubois, and Ohler, 2014, Alytes, 30: 25–26, and supported by Nečas, Kielgast, Chinemerem, Rödel, Dolinay, and Gvoždík, 2024, Zool. J. Linn. Soc., 202(4): Suppl. Material.

Hyperolius maximus Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 91. Holotype: ZMB unnumbered, according to the original publication; this being ZMB 36113, according to Tillack, de Ruiter, and Rödel, 2021, Zoosyst. Evol., Berlin, 97: 428; MCZ 17639 considered 'cotype', on exchange from ZMB by Barbour and Loveridge, 1946, Bull. Mus. Comp. Zool., 96: 127; presumably this is a paratype (DRF). Type locality: "Ossindinge", Cameroon. Synonymy by Loveridge, 1938, Proc. New England Zool. Club, 17: 49–74; Laurent, 1961, Rev. Zool. Bot. Afr., 64: 78. The description in,Ahl, 1931, Das Tierreich, 55: 366, appeared subsequently according to Tillack, de Ruiter, and Rödel, 2021, Zoosyst. Evol., Berlin, 97: 428.

Hyperolius hildebrandti Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 101. Holotype: ZMB unnumbered, according to the original publication; ZMB 8378, according to Laurent, 1961, Rev. Zool. Bot. Afr., 64: 73, and Tillack, de Ruiter, and Rödel, 2021, Zoosyst. Evol., Berlin, 97: 425. Type locality: "Kamerun"; Doula Region, Cameroon, according to Tillack, de Ruiter, and Rödel, 2021, Zoosyst. Evol., Berlin, 97: 425. Synonymy with Hyperolius guttatus by Frétey, Dubois, and Ohler, 2014, Alytes, 30: 20.

Hyperolius ocellatus hildebrandti — Laurent, 1950, Explor. Parc Natl. Albert, Miss. G.F. de Witte (1933–1935), 64: 107; Laurent, 1951, Ann. Soc. R. Zool. Belg., 82: 48; Perret and Mertens, 1957, Bull. Inst. Franç. Afr. Noire, Ser. A, 19: 571. Distinctiveness from Hyperolius ocellatus purpurescens rejected by Perret, 1975, Ann. Fac. Sci. Cameroun, 20: 23–31.

Hyperolius ocellatus guttatus — Laurent, 1956, Rev. Zool. Bot. Afr., 53: 235; Laurent, 1958, Mem. Inst. Franç. Afr. Noire, 53: 288;Perret, 1966, Zool. Jahrb., Jena, Abt. Syst., 93: 401; Schiøtz, 1966, Vidensk. Medd. Dansk Naturhist. Foren., 129: 48. See Perret, 1975, Ann. Fac. Sci. Cameroun, 20: 23–31, and Lötters, Gossmann, Obame, and Böhme, 2001, Herpetofauna, Weinstadt, 23: 29–30.

Hyperolius concolor guttatus — Frétey, Dubois, and Ohler, 2014, Alytes, 30: 20.

Hyperolius guttatus — Nečas, Kielgast, Chinemerem, Rödel, Dolinay, and Gvoždík, 2024, Zool. J. Linn. Soc., 202(4): 13 (Zoobank Publication registration: 004579A5-F55C-4429-97C6-A52A23F4501D, 15 May 2024)

Common Names

None noted.

Distribution

Southwestern Cameroon, possibly coastal south-eastern Nigeria east of the Cross River.

Geographic Occurrence

Natural Resident: Cameroon

Likely/Controversially Present: Nigeria

Endemic: Cameroon

Comment

Resurrected on the basis of molecular markers from the synonymy of Hyperolius concolor by Nečas, Kielgast, Chinemerem, Rödel, Dolinay, and Gvoždík, 2024, Zool. J. Linn. Soc., 202(4): 1–19, who discussed systematics, morphology, molecular markers, and distribution, including a dot map.

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