Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hyperolius nasutus Günther, 1865

Class: Amphibia > Order: Anura > Family: Hyperoliidae > Subfamily: Hyperoliinae > Genus: Hyperolius > Species: Hyperolius nasutus

Hyperolius nasutus Günther, 1865 "1864", Proc. Zool. Soc. London, 1864: 482. Holotype: BMNH 1947.2.9.68 (formerly 1864.10.28.14), by museum records. Type locality: "Duque de Bragança", Angola.

Rappia nasuta — Günther, 1869 "1868", Proc. Zool. Soc. London, 1868: 481; Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 127.

Rappia punctulata Barboza du Bocage, 1895, Herpetol. Angola Congo: 168. Holotype: MBL T.23-227 according to Perret, 1976, Arq. Mus. Bocage, Ser. 2, 6: 27; destroyed in 1978 fire according to Laurent, 1985, in Frost (ed.), Amph. Species World: 216. Type locality: "sur les bords du Quanza", Angola. Synonymy by Channing, Hillers, Lötters, Rödel, Schick, Conradie, Rödder, Mercurio, Wagner, Dehling, Du Preez, Kielgast, and Burger, 2013, Zootaxa, 3620: 334.

Hyperolius punctulatus — Noble, 1924, Bull. Am. Mus. Nat. Hist., 49: 251, 328; Ahl, 1931, Das Tierreich, 55: 277.

Hyperolius nasutus — Noble, 1924, Bull. Am. Mus. Nat. Hist., 49: 251, 327.

Hyperolius nasutus nasutus — Laurent, 1954, Publ. Cult. Companhia Diamantes Angola, 23: 136 (17 of preprint); De Witte, 1952, Result. Sci., Explor. Hydrobiol. Lac Tanganyika (1946–47), 3: 12; Laurent, 1957, Explor. Parc Natl. Upemba, Miss. G.F. de Witte, 42: 27; Poynton, 1964, Ann. Natal Mus., 17 192.

English Names

Large-nosed Long Reed Frog (Channing, Hillers, Lötters, Rödel, Schick, Conradie, Rödder, Mercurio, Wagner, Dehling, Du Preez, Kielgast, and Burger, 2013, Zootaxa, 3620: 334).

Longnose Reed Frog (Stewart and Wilson, 1966, Ann. Natal Mus., 18: 304; Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 67).

Long-nosed Reed Frog (Van Dijk, 1978, J. Herpetol. Assoc. Afr., 17: 16; Ananjeva, Borkin, Darevsky, and Orlov, 1988, Dict. Amph. Rept. Five Languages: 73).

Sharp-nosed Reed Frog (Ananjeva, Borkin, Darevsky, and Orlov, 1988, Dict. Amph. Rept. Five Languages: 73).

Sharp-and-blunt-snouted Sedge Frog (Ananjeva, Borkin, Darevsky, and Orlov, 1988, Dict. Amph. Rept. Five Languages: 73).

Long Reed Frog (Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 254).

Quanza Reed Frog (Hyperolius punctulatus [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 68).

Large-nosed Reed Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 176).

Distribution

Southern Africa from northern Angola and the Okovango Delta of Botswana; presumably in intervening Caprivi Strip of Namibia and likely into western Zambia. See comment.

Comment

Schiøtz, 1999, Treefrogs Afr.: 97–100, provided an account for the species in the sense of including Hyperolius granulatus and Hyperolius benguellensis as synonyms. Channing, Moyer, and Burger, 2002, Afr. Zool., 37: 91–99, partitioned nominal Hyperolius nasutus into three cryptic species (Hyperolius nasutus, Hyperolius viridis, and Hyperolius acuticeps [now a synonym of Hyperolius microps]) on the basis of call-type and morphometrics. They retained Hyperolius benguellensis in the synonymy of Hyperolius nasutus. Schiøtz and Van Daele, 2003, Alytes, 20: 138–142, distinguished two species in this complex in Angola on the basis of call structure and color pattern, one of which they assigned the name Hyperolius benguellensis and the other to Hyperolius nasutus. Although their evidence clearly supports two taxa existing within nominal Hyperolius nasutus, it is not clear that the names are applied appropriately given that types were not examined and the possibility of unnamed members of this complex being involved did not seem to be addressed. Nago, Grell, Sinsin, and Rödel, 2006, Salamandra, 42: 93–108, provided a record of nominal Hyperolius nasutus for Benin. Discussed (as Hyperolius punctulatus) by Perret, 1976, Arq. Mus. Bocage, Ser. 2, 6: 27. Channing, Hillers, Lötters, Rödel, Schick, Conradie, Rödder, Mercurio, Wagner, Dehling, Du Preez, Kielgast, and Burger, 2013, Zootaxa, 3620: 334–335, provided an account. Harper, Measey, Patrick, Menegon, and Vonesh, 2010, Field Guide Amph. E. Arc Mts. Tanzania and Kenya: 174–175, provided a brief account of nominal Hyperolius nasutus, but inasmuch as the species is extralimital to the area of their guide it is unclear what species is being addressed. Marques, Ceríaco, Blackburn, and Bauer, 2018, Proc. California Acad. Sci., Ser. 4, 65 (Suppl. II): 106–108, provided a map for Angola and a brief account. Butler, Ceríaco, Marques, Bandeira, Júlio, Heinicke, and Bauer, 2019, Herpetol. Rev., 50: 225–240, reported on a collection from Huila Province, southwestern Angola, as Hyperolius cf. nasutus. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 176–177, provided a brief account, photograph, and range map. Mali, Banda, Chifundera, Badjedjea, Sebe, Lokasola, Ewango, Tungaluna, and Akaibe, 2019, Am. J. Zool., 2: 38–43, reported the species from the Okapi Wildlife Reserve, northeastern Dem. Rep. Congo, although the identification of this is now suspect.

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