Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hyperolius papyri (Werner, 1908 "1907")

Class: Amphibia > Order: Anura > Family: Hyperoliidae > Subfamily: Hyperoliinae > Genus: Hyperolius > Species: Hyperolius papyri

Rappia papyri Werner, 1908 "1907", Sitzungsber. Akad. Wiss. Wien, Phys. Math. Naturwiss. Kl., 116: 1903. Holotype: NHMW 14847, according to Häupl and Tiedemann, 1978, Kat. Wiss. Samml. Naturhist. Mus. Wien, 2: 28, and Häupl, Tiedemann, and Grillitsch, 1994, Kat. Wiss. Samml. Naturhist. Mus. Wien, 9: 33. Type locality: "Khor Attar und Mongalla . . . Gondokoro"; locality associated with holotype is Gondoroko, South Sudan.

Rappia pachyderma Werner, 1908 "1907", Sitzungsber. Akad. Wiss. Wien, Phys. Math. Naturwiss. Kl., 116: 1903. Holotype: NHMW 14847, according to Häupl and Tiedemann, 1978, Kat. Wiss. Samml. Naturhist. Mus. Wien, 2: 28, and Häupl, Tiedemann, and Grillitsch, 1994, Kat. Wiss. Samml. Naturhist. Mus. Wien, 9: 34. Type locality: "Steppe von Gondoroko", Uganda. Considered a subspecies of Hyperolius viridiflavus by Laurent, 1951, Ann. Soc. R. Zool. Belg., 82: 40; Laurent, 1961, Rev. Zool. Bot. Afr., 64: 92, who did not examine the types and confused this species with the Hyperolius viridiflavus complex. Synonymy by Dehling, 2015, Herpetozoa, Wien, 28: 39.

Hyperolius pachydermus — Noble, 1924, Bull. Am. Mus. Nat. Hist., 49: 252, 327.

Hyperolius pachyderma — Ahl, 1931, Das Tierreich, 55: 386. Incorrect subsequent spelling.

Hyperolius viridiflavus pachyderma — Laurent, 1951, Ann. Soc. R. Zool. Belg., 82: 40.

Hyperolius viridiflavus pachydermus — Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 392. Incorrect subsequent spelling based on specimens of the Hyperolius viridiflavus complex.

Hyperolius papyri — Dehling, 2015, Herpetozoa, Wien, 28: 39.

Common Names

Nile Reed Frog (Spawls, Mazuch, and Mohammad, 2023, Handb. Amph. Rept. NE Afr.: 70).

Distribution

Woodland and savannah in the upper reaches of the White Nile in Sudan, most of eastern South Sudan, and Ethiopia west of the Rift Valley, 200 to ca. 2000 m elevation.

Geographic Occurrence

Natural Resident: Ethiopia, South Sudan, Sudan

Comment

Hyperolius papyri had been placed in the synonymy of Hyperolius nasutus by Noble, 1924, Bull. Am. Mus. Nat. Hist., 49: 259, and Schiøtz, 1967, Spolia Zool. Mus. Haun., 25: 139. See discussion by Channing, Moyer, and Burger, 2002, Afr. Zool., 37: 97. In the Hyperolius nasutus group of Amiet, 2005, Rev. Suisse Zool., 112: 271–310, and recognized as a distinct species by Dehling, 2015, Herpetozoa, Wien, 28: 39–47, who examined the relevant types and discussed the history of the use, and abuse, of the name and various synonymies. Spawls, Mazuch, and Mohammad, 2023, Handb. Amph. Rept. NE Afr.: 69–70, provided an account addressing identification, natural history, conservation status, and range, including a polygon map.

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