Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hyperolius thomensis Barboza du Bocage, 1886

Class: Amphibia > Order: Anura > Family: Hyperoliidae > Subfamily: Hyperoliinae > Genus: Hyperolius > Species: Hyperolius thomensis

Hyperolius thomensis Barboza du Bocage, 1886, J. Sci. Math. Phys. Nat., Lisboa, 11 (42): 74. Syntypes: MBL T.26-216 (2 specimens) and 26-219; destroyed by 1978 fire. CAS 251637 designate neotype by Bell, 2016, Herpetologica, 72: 349. Type locality: "St. Thomé"; corrected to "Roça Saudade e littoral", São Thomé Island by Barboza du Bocage, 1897, J. Sci. Math. Phys. Nat., Lisboa, Ser. 2, 6: 203. Neotype from "São Tomé and Príncipe, São Tomé, trail from Bom Sucesso Radio Tower, 0.275688 N, 6.604128 E, 1326 m elevation".

Rappia thomensis — Boulenger, 1887, Zool. Rec., 23: 21, by implication.

Nesionixalus thomensis — Perret, 1976, Arq. Mus. Bocage, Ser. 2, 6: 29.

Hyperolius (Nesionixalus) thomensis — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 366.

Common Names

São Tomé Reed Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 192).

Distribution

São Tomé Island in primary forest, 300 to 1300 m elevation, São Tomé I., Gulf of Guinea.

Geographic Occurrence

Natural Resident: São Tomé and Principe

Endemic: São Tomé and Principe

Comment

See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 288. Schiøtz, 1999, Treefrogs Afr.: 311, provided a brief account and map as Nesionixalus thomensis. Biogeography and diversification reported on by Bell, Drewes, Channing, Gvoždík, Kielgast, Lötters, Stuart, and Zamudio, 2015, J. Biogeograph., 42: 65–75, and Bell, Drewes, and Zamudio, 2015, Evolution, 69: 904–915, suggesting that Hyperolius molleri (now partitioned into Hyperolius molleri and Hyperolius drewesi) is the sister taxon of this species, and together the sister taxon of Hyperolius cinnamomeoventris. Drewes and Stoelting, 2004, Proc. California Acad. Sci., Ser. 4, 55: 574–575, provided life history observations and discussed problems with published collection locations. Gilbert and Bell, 2017 "2018", Biol. J. Linn. Soc., 123: 1–11, compared call structure of the Hyperolius thomenssi/Hyperolius moleri/Hyperolius drewesi/Hyperolius olivaceus group in São Tome and Principe and Central Africa. Dehling and Sinsch, 2019, Zool. Anz., 280: 65–77, included this species in their Hyperolius cinnamomeoventris group. Bell and Irian, 2019, Biol. J. Linn. Soc., 128: 672–680, reported on a hybrid zone with Hyperolius molleri. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 192–193, provided a brief account, photograph, and range map. Bell, Ceríaco, Scheinberg, and Drewes, 2022, in Ceríaco et al. (eds.), Biodiversity Gulf Guinea Islands: 491–494, summarized the literature and systematics of the species. Ceríaco, Marques, Sousa, Veríssimo, Beja, and Ferreira, 2023, ZooKeys, 1168: 41–75, included this species in a report, key, and DNA barcode reference library for the amphibians of São Tomé and Príncipe.

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

For access to general information see Wikipedia
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For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.