Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Boophis tephraeomystax (Duméril, 1853)

Class: Amphibia > Order: Anura > Family: Mantellidae > Subfamily: Boophinae > Genus: Boophis > Species: Boophis tephraeomystax

Polypedates tephraeomystax Duméril, 1853, Ann. Sci. Nat., Paris, Ser. 3, 19: 158. Syntypes: MNHNP 4584 (2 specimens, according to Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 53), MNHNP 1660A-B and 4584 (according to Guibé, 1978, Bonn. Zool. Monogr., 11: 65), and NHMW 22880 (on exchange from MNHNP), according to Häupl and Tiedemann, 1978, Kat. Wiss. Samml. Naturhist. Mus. Wien, 2: 26, and Häupl, Tiedemann, and Grillitsch, 1994, Kat. Wiss. Samml. Naturhist. Mus. Wien, 9: 31. MNHNP 1994.1040 designated lectotype by Glaw and Vences, 1997, Copeia, 1997: 576. Type locality: "Madagascar".

Polypedates crossleyi Peters, 1874, Monatsber. Preuss. Akad. Wiss. Berlin, 1874: 618. Holotype: ZMB 8216 according to Bauer, Günther, and Klipfel, 1995, in Bauer et al. (eds.), Herpetol. Contr. W.C.H. Peters: 51. Type locality: "Innern von Madagascar (Nossi Vola)". Synonymy by Guibé, 1978, Bonn. Zool. Monogr., 11: 65.

Polypedates dispar Boettger, 1878, Ber. Senckenb. Naturforsch. Ges., 1878–79: 86. Types: SMF; SMF 6766 designated lectotype by Mertens, 1967, Senckenb. Biol., 48(A): 48. Type locality: "Insula Nossi-Bé", Madgascar. Synonymy by Boulenger, 1889, Ann. Mag. Nat. Hist., Ser. 6, 4: 246; Ahl, 1931, Das Tierreich, 55: 200; Blommers-Schlösser, 1979, Bijdr. Dierkd., 49: 297.

Polypedates dispar var. leucopleura Boettger, 1881, Zool. Anz., 4: 47. Holotype: Bremen Museum, by original designation; presumably lost. Type locality: "insular Madagascar"; restricted to Nosy Bé by Boettger, 1882, Abh. Naturwiss. Ver. Bremen, 7: 177-190..

Rhacophorus crossleyi — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 77.

Rhacophorus dispar — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 77.

Rhacophorus tephraeomystax — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 78.

Rhacophorus difficilis Boettger, 1892, Kat. Batr. Samml. Mus. Senckenb. Naturforsch. Ges.: 14. Holotype: SMF 6762 (formerly 1075a) according to Mertens, 1967, Senckenb. Biol., 48(A): 48. Type locality: "Foizana, O. Madagascar". Synonymy by Guibé, 1978, Bonn. Zool. Monogr., 11: 65; the synonymy rejected by Blommers-Schlösser, 1979, Bijdr. Dierkd., 49: 261; but reaffirmed by Glaw, Vences, Andreone, and Vallan, 2001, Zool. J. Linn. Soc., 133: 495.

Rhacophorus hildebrandti Ahl, 1925, Zool. Anz., 65: 18. Holotype: ZMB unnumbered in the original publication; given as ZMB 10472 by Guibé, 1978, Bonn. Zool. Monogr., 11: 65, and Blommers-Schlösser, 1979, Bijdr. Dierkd., 49: 297. Type locality: "Nordwest-Madagaskar". Synonymy by Guibé, 1978, Bonn. Zool. Monogr., 11: 65; Blommers-Schlösser, 1979, Bijdr. Dierkd., 49: 297.

Rhacophorus (Rhacophorus) hildebrandti — Ahl, 1931, Das Tierreich, 55: 199.

Rhacophorus (Rhacophorus) difficilis — Ahl, 1931, Das Tierreich, 55: 202.

Rhacophorus (Rhacophorus) tephraeomystax — Ahl, 1931, Das Tierreich, 55: 200.

Rhacophorus (Rhacophorus) crossleyi — Ahl, 1931, Das Tierreich, 55: 202.

Boophis crossleyi — Guibé, 1947, Bull. Mus. Natl. Hist. Nat. Paris, Ser. 2, 19: 439.

Boophis difficilis — Guibé, 1947, Bull. Mus. Natl. Hist. Nat. Paris, Ser. 2, 19: 439. Blommers-Schlösser, 1979, Bijdr. Dierkd., 49: 261; Glaw and Vences, 1992, Fieldguide Amph. Rept. Madagascar: 280.

Rhacophorus (Boophis) crossleyi — Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 52.

Rhacophorus (Boophis) tephraeomystax — Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 53.

Boophis tephraeomystax — Blommers-Schlösser, 1979, Bijdr. Dierkd., 49: 296.

Boophis (Sahona) tephaeomystax — Glaw and Vences, 2006, Organisms Divers. Evol., 6: 242, by implication; Glaw and Vences, 2006, Organisms Divers. Evol., Electron. Suppl., 11(1): 1.

Common Names

Dumeril's Bright-eyed Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 111).

Foizana Bright-eyed Frog (Boophis difficilis [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 110).

Distribution

Eastern and northwestern Madagascar; Nosy Bé and adjacent Nosy Komba.

Geographic Occurrence

Natural Resident: Madagascar

Endemic: Madagascar

Comment

In the Boophis tephraeomystax group, according to Blommers-Schlösser, 1979, Bijdr. Dierkd., 49: 296. In the Boophis tephraeomystax group of Glaw and Vences, 2006, Organisms Divers. Evol., Electron. Suppl., 11(1): 1. See discussion by Glaw, Vences, Andreone, and Vallan, 2001, Zool. J. Linn. Soc., 133: 495-529. See comment under Boophis doulioti. Andreone, Glaw, Nussbaum, Raxworthy, Vences, and Randrianirina, 2003, J. Nat. Hist., London, 37: 2119-2149, discussed the occurrence of this species on Nosy Be. Glaw and Vences, 2007, Field Guide Amph. Rept. Madagascar, Ed. 3: 146-147, provided an account. Roberts and Daly, 2014, Salamandra, 50: 18–26, reported this species from Nosy Komba, between Nosy Be and the mainland. Rodriguez, Börner, Pabijan, Gehara, Haddad, and Vences, 2015, Evol. Ecol., 29: 765–785 (and supplemental data), reported on phylogeographic diversity and its causes.

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