Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Mantidactylus guttulatus (Boulenger, 1881)

Class: Amphibia > Order: Anura > Family: Mantellidae > Subfamily: Mantellinae > Genus: Mantidactylus > Species: Mantidactylus guttulatus

Rana guttulata Boulenger, 1881, Ann. Mag. Nat. Hist., Ser. 5, 7: 360. Syntypes: BMNH 1947.2.25.48–52 (formerly 1880.7.15.10–12, 1881.3.29.15–16 according to Guibé, 1978, Bonn. Zool. Monogr., 11: 18); BMNH 1947.2.25.51 designated lectotype by Blommers-Schlösser and Blanc, 1991, Faune de Madagascar, 75: 256. Type locality: "Betsileo", Madagascar ; corrected to "S.E. Betsileo", Madagascar, by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 21.

Mantidactylus guttulatus — Boulenger, 1895, Ann. Mag. Nat. Hist., Ser. 6, 15: 450.

Mantidactylus (Mantidactylus) guttulatus — Ahl, 1931, Das Tierreich, 55: 12; Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 313; Glaw and Vences, 2006, Organisms Divers. Evol., Electron. Suppl., 11(1): 3.

English Names

Gray Madagascar Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 101).

Distribution

Central part of eastern Madagascar.

Geographics occurrence

Natural resident: Madagascar

Endemic to the political unit: Madagascar

Comment

Glaw and Vences, 2007, Field Guide Amph. Rept. Madagascar, Ed. 3 230–231, provided an account. Altig and McDiarmid, 2006, Proc. Biol. Soc. Washington, 119: 418–425, described larval morphology. Schulze, Randrianiaina, Perl, Glaw, and Vences, 2016, Acta Herpetol., Firenze, 11: 119–125, reported on larval morphology. Rodriguez, Börner, Pabijan, Gehara, Haddad, and Vences, 2015, Evol. Ecol., 29: 765–785 (and supplemental data), reported on phylogeographic diversity and its causes. Rancilhac, Bruy, Scherz, Pereira Silva, Preick, Straube, Lyra, Ohler, Streicher, Andreone, Crottini, Hutter, Randrianantoandro, Rakotoarison, Glaw, Hofreiter, and Vences, 2020, J. Nat. Hist., London, 54: 87–118, discussed the systematics and the range.

External links:

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For additional sources of information from other sites search Google
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.