Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptobrachium liui (Pope, 1947)

Class: Amphibia > Order: Anura > Family: Megophryidae > Subfamily: Leptobrachiinae > Genus: Leptobrachium > Species: Leptobrachium liui

Vibrissaphora liui Pope, 1947, Copeia, 1947: 109. Holotype: FMNH 24427, by original designation. Type locality: "region of San Chiang [= Sangang], Ch'ungan Hsien [now Wuyishan City], Fukien [=Fujian] Province, China".

Vibrissaphora yaoshanensis Liu and Hu in Liu, Hu, Tian, and Wu, 1978, Mater. Herpetol. Res., 4: 18. Holotype: CIB 61001, by original designation. Type locality: Yangliuchong, Jinxiu, Yaoshan, Guangxi, altitude 1200 m, China. Synonymy by Wu and Zhao, 1987, Acta Herpetol. Sinica, Chengdu, N.S.,, 6 (3): 42–44; Fei, Ye, Jiang, Chen, Cai, Tang, Cai, Gao, and Wei, 1995, Acta Herpetol. Sinica, 4–5: 68; Dubois and Ohler, 1998, Dumerilia, 4: 27, and Zheng, Li, and Fu, 2008, Mol. Phylogenet. Evol., 46: 704.

Leptobrachium (Leptobrachium) liui — Dubois, 1980, Bull. Mens. Soc. Linn. Lyon, 49: 476. .

Vibrissaphora liui — Liu, Hu, and Zhao, 1980, Acta Herpetol. Sinica, 3 (1): 3; Zhao and Adler, 1993, Herpetol. China: 126.

Vibrissaphora jiulongshanensis Wei and Zhao, 1981, J. Hangzhou Univ., Nat. Sci., 8: 300. Types: HU 800013, by original designation. Type locality: "Jiulongshan of Shuichangxian, Zhejiang [Province], 900 m", China. Synonymy by Huang, Gu, and Zong, 1984, J. Hangzhou Normal College (Nat. Hist.), 1984: 14. Synonymy confirmed by Zheng, Li, and Fu, 2008, Mol. Phylogenet. Evol., 46: 704.

Vibrissaphora yaoshanensis — Ye, Fei, and Hu, 1993, Rare and Economic Amph. China: 156.

Vibrissaphora liui yaoshanensis — Fei, Ye, Jiang, Chen, Cai, Tang, Cai, Gao, and Wei, 1995, Acta Herpetol. Sinica, 4–5: 68.

Vibrissaphora liui liui — Fei, Ye, Jiang, Chen, Cai, Tang, Cai, Gao, and Wei, 1995, Acta Herpetol. Sinica, 4–5: 68.

Leptobrachium (Vibrissaphora) liui liui — Dubois and Ohler, 1998, Dumerilia, 4: 27.

Leptobrachium (Vibrissaphora) liui yaoshanense — Dubois and Ohler, 1998, Dumerilia, 4: 27.

Vibrissaphora yaoshanensis — Ho, Lathrop, Murphy, and Orlov, 1999, Russ. J. Herpetol., 6: 49; Rao and Wilkinson, 2008, Mol. Phylogenet. Evol., 46: 68–69.

Leptobrachium (Vibrissaphora) liui — Delorme, Dubois, Grosjean, and Ohler, 2006, Alytes, 24: 12; Matsui, Hamidy, Murphy, Khonsue, Imbun, Shimada, Ahmad, Belabut, and Jiang, 2010, Mol. Phylogenet. Evol., 56: 269.

Leptobrachium liui — Rao and Wilkinson, 2008, Mol. Phylogenet. Evol., 46: 69.

Leptobrachium yaoshanense — Rao and Wilkinson, 2008, Mol. Phylogenet. Evol., 46: 69.

Leptobrachium liui yaoshanensis — Shen, 2014, Fauna Hunan, Amph.: 136. Incorrect subsequent spelling of yaoshanense.

Common Names

Pope's Spiny Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 87).

Chong'an Moustache Toad (Fei, 1999, Atlas Amph. China: 96).

崇安髭蟾 (Chinese: Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang, and Che, 2020, Biodiversity Sci., 28: App. 1: 4).

Yaoshan Moustache Toad (Leptobrachium yaoshanense: Fei, 1999, Atlas Amph. China: 98).

Distribution

Northern Guangxi, northern Guangdong, southern and northern Hunan, central Jiangxi, southern Zhejiang, northern Fujian, and northern Guangdong, China.

Geographic Occurrence

Natural Resident: China, People's Republic of

Endemic: China, People's Republic of

Comment

Reviewed (as Vibrissaphora liui) by Liu, Hu, and Zhao, 1980, Acta Herpetol. Sinica, 3 (1): 1–9. See Pope, 1931, Bull. Am. Mus. Nat. Hist., 61: 447–450 (as Vibrissaphora pelodytoides). Pope, 1947, Copeia, 1947: 109–112 (as Vibrissaphora liui). See accounts by Ye, Fei, and Hu, 1993, Rare and Economic Amph. China 155; Dubois and Ohler, 1998, Dumerilia, 4: 27; and Fei, 1999, Atlas Amph. China: 98–99. See comment under Leptobrachium hasseltii. Huang, 1990, Fauna Zhejiang, Amph. Rept.: 40–43, provided an account (as Vibrissaphora liui) for Zhejiang populations. In the Vibrissaphora liui group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 70. Reviewed (as Vibrissaphora yaoshanensis) by Liu, Hu, and Zhao, 1980, Acta Herpetol. Sinica, 3 (1): 1–9. The status of Leptobrachium yaoshanense is arguable at this point in time (2008, DRF) with molecular evidence arguing for both species boundaries (e.g., Rao and Wilkinson, 2008, Mol. Phylogenet. Evol., 46: 61–73) and the lack thereof (e.g., Zheng, Li, and Fu, 2008, Mol. Phylogenet. Evol., 46: 695–707). Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 2: 286–297, provided accounts for the two nominal subspecies, spot maps, and illustrations. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 165–166, provided brief accounts (as Vibrissaphora liu liu, page 165; and Vibrissaphora liui yaoshanensis) including photographs of specimens and habitat. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 193–195, provided accounts (as Vibrissaphora liui liui and Vibrissaphora liui yaoshanensis), photographs, and a range map for China. Fei and Ye, 2016, Amph. China, 1: 480–486, provided account (as Vibrissaphora liui liui and Vibrissaphora liui yaoshanensis), photographs, and dot maps. Shen, 2014, Fauna Hunan, Amph.: 136–140, provided an account for Hunan. China. Li, 2011, Amph. Rept. Guangdong: 22, provided a brief account for Guangdong, China, and photograph. Ding, Chen, Tang, Zheng, and Ji, 2020, Zootaxa, 4732: 585–588, reported on the advertisement call. Li, Ai, Xie, Huang, Qiu, Fu, Zhao, Fu, and Wu, 2024, Mol. Ecol., 33(e17569): 1–18, discussed ancient introgression of Leptobrachium leishanensis into Leptobrachium liui, and provided a molecular tree of phylogenetic placement and a dot and polygon map of the distribution. Hu, Feng, Zhang, Xiang, Hao, Zhou, and Ding, 2025, Asian Herpetol. Res., 16: 192–201, discussed advertisement call variability in the Zhejiang Jiulongshan National Nature Reserve, Suichang County, Zhejiang Province, China. Zhang, Zhang, Wu, Tang, Sheng, Xie, Huang, Ou, Xiao, and Xiang, 2026, Life Sci. Res., 30: 124–136, discussed habitat in Zhangjiajie National Forest Park, Hunan Province, China.

External links:

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China