Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Boulenophrys tuberogranulata (Shen, Mo and Li, 2010)

Class: Amphibia > Order: Anura > Family: Megophryidae > Subfamily: Megophryinae > Genus: Boulenophrys > Species: Boulenophrys tuberogranulata

Megophrys tuberogranulatus Shen, Mo and Li In Mo, Shen, Li, and Wu, 2010, Curr. Zool., Chengdu, 56: 433. Holotype: HNUL03080902, by original designation. Type locality: "Tianzishan Nature Reserve, Sangzhi County, Hunan Province (29°20′-29°50′N, 110°20′-110°30′E), elevation 1130 m". Gender disagreement of genus and species name.

Xenophrys tuberogranulatus — Chen, Zhou, Poyarkov, Stuart, Brown, Lathrop, Wang, Yuan, Jiang, Hou, Chen, Suwannapoom, Nguyen, Duong, Papenfuss, Murphy, Zhang, and Che, 2017, Mol. Phylogenet. Evol., 106: 41.

Megophrys tuberogrannulus — Shen, 2014, Fauna Hunan, Amph.: 171. Incorrect subsequent spelling.

Boulenophrys tuberogranulatus — Fei and Ye, 2016, Amph. China, 1: 663. Gender disagreement.

Xenophrys (Panophrys) tuberogranulatus — Chen, Zhou, Poyarkov, Stuart, Brown, Lathrop, Wang, Yuan, Jiang, Hou, Chen, Suwannapoom, Nguyen, Duong, Papenfuss, Murphy, Zhang, and Che, 2017, Mol. Phylogenet. Evol., 109: 41.

Megophrys (Panophrys) tuberogranulatus — Mahony, Foley, Biju, and Teeling, 2017, Mol. Biol. Evol., 34: 755.

Panophrys tuberogranulata — Lyu, Zeng, Wang, Liu, Huang, Li, and Wang, 2021, Zootaxa, 4927: 13. Correction in gender of species name.

Boulenophrys tuberogranulata — Qi, Lyu, Wang, Mo, Zeng, Zeng, Dai, Li, Grismer, and Wang, 2021, Zootaxa, 5072: 404.

Common Names

Tianzishan Horned Toad (Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 241).

Distribution

Known only from Tianzishan Nature Reserve and Mount Tianping in northwestern Hunan, China, at elevations of 1000–1380 m.

Geographic Occurrence

Natural Resident: China, People's Republic of

Endemic: China, People's Republic of

Comment

Most closely related to Megophrys kuatunensis and Megophrys minor according to the original publication. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 241, provided an account (as Megophrys tuberogranulatus), photographs, and a range map. Fei and Ye, 2016, Amph. China, 1: 663–665, provided an account, photographs, and dot map. Shen, 2014, Fauna Hunan, Amph.: 171–174, provided a detailed account for Hunan, China. Lyu, Qi, Wang, Zhang, Zhao, Zeng, Wan, Yang, Mo, and Wang, 2023, Zool. Res., Kunming, 44: 440, provided an account and placed this in their Boulenophrys boettgeri group. Xiang, Zhou, Wei, Shu, Gu, and Jiang, 2024, Ecol. Evol., 4(e11687): 1–13, reported on the complete mitogenome.

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China