Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Rhombophryne coudreaui (Angel, 1938)

Class: Amphibia > Order: Anura > Family: Microhylidae > Subfamily: Cophylinae > Genus: Rhombophryne > Species: Rhombophryne coudreaui

Plethodontohyla coudreaui Angel, 1938, Bull. Mus. Natl. Hist. Nat. Paris, Ser. 2, 10: 260. Holotype: MNHNP 1937.19, according to Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 61. Type locality: "Reeserve de Betampona (forêt orientale, près de Moramanga, à mi-chemin, entre Tamatave et Tananarive)", Madagascar.

Rhombophryne coudreaui — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 366.

Common Names

Betampona Digging Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 92).

Distribution

Low-elevation forests in eastern Madagascar, and is with certainty known only from (1) Réserve Naturelle Intégrale de Betampona (type locality), where it has been found at Sahambendrana, Maintimbato and Sahaindrana as well as at Sahabefoza (17.9142°S, 49.2078°E, 353 m elevation); photographic evidence from Ampasina, a highly degraded forest just 8 km east of Betampona (ca 80 m elevation). Elevation: ca 80–200 m

Geographic Occurrence

Natural Resident: Madagascar

Endemic: Madagascar

Comment

Andreone, 2003, Herpetol. Rev., 34: 259, detailed the known localities of this frog. See brief account by Glaw and Vences, 2007, Field Guide Amph. Rept. Madagascar, Ed. 3: 118–119. See photograph, map, description of geographic range and habitat, and conservation status (as Plethodontohyla coudreaui) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 457. In the Rhombophryne testudo species group of Belluardo, Scherz, Santos, Andreone, Antonelli, Glaw, Muñoz-Pajares, Randrianirina, Raselimanana, Vences, and Crottini, 2022, Syst. Biodiversity, 20 (1: 2039320): 1–13. Scherz, Glaw, Petzold, Andreone, Köhler, Raselimanana, Hutter, Belluardo, Rakotoarison, Hofreiter, Vences, and Crottini, 2026, Vert. Zool., Senckenberg, 76: 485–610, provided a species account covering phylogenetic relationships, morphology, osteology, natural history, and a corrected statements of distribution and identification.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observations see iNaturalist